ratio of word probabilities predicted from brain for ant and bicycle

ant	bicycle
top 10 words in brain distribution (in article): engine energy gas cell produce air speed drive type increase	top 10 words in brain distribution (in article): city build power male station human state common time train
top 10 words in brain distribution (not in article): vehicle fuel wheel power car gear design aircraft oil hydrogen	top 10 words in brain distribution (not in article): animal species cat breed wolf hunt dog wild population bear
times more probable under ant 30 20 10 6 4 2.5 1.25 1 1.25 2.5 4 6 10 20 30 times more probable under bicycle (words not in the model)
A phylogeny of the extant ant subfamilies. "'Ants'" are social insects of the family "'Formicidae'", and along with the related wasps and bees, they belong to the order Hymenoptera. Ants evolved from wasp-like ancestors in the mid-Cretaceous period between 110 and 130 million years ago and diversified after the rise of flowering plants. Today, more than 12,000 species are classified with upper estimates of about 14,000 species. They are easily identified by their elbowed antennae and a distinctive node-like structure that forms a slender waist. Ants form colonies that range in size from a few dozen predatory individuals living in small natural cavities to highly organised colonies which may occupy large territories and consist of millions of individuals. These larger colonies consist mostly of sterile wingless females forming castes of "workers", "soldiers", or other specialised groups. Ant colonies also have some fertile males called "drones" and one or more fertile females called "queens". The colonies are sometimes described as superorganisms because ants appear to operate as a unified entity, collectively working together to support the colony. Ants have colonised almost every landmass on Earth. The only places lacking indigenous ants are Antarctica and certain remote or inhospitable islands. Ants thrive in most ecosystems, and may form 15–25% of the terrestrial animal biomass. Their success has been attributed to their social organisation and their ability to modify habitats, tap resources, and defend themselves. Their long co-evolution with other species has led to mimetic, commensal, parasitic, and mutualistic relationships. Ant societies have division of labour, communication between individuals, and an ability to solve complex problems. These parallels with human societies have long been an inspiration and subject of study. Many human cultures make use of ants in cuisine, medication and rituals. Some species are valued in their role as biological pest control agents. However, their ability to exploit resources brings ants into conflict with humans, as they can damage crops and invade buildings. Some species, such as the red imported fire ant, are regarded as invasive species, since they have establish themselves in new areas where they may be accidentally introduced. Taxonomy and evolution. The family Formicidae belongs to the order Hymenoptera, which also includes sawflies, bees and wasps. Ants evolved from a lineage within the vespoid wasps. Phylogenetic analysis suggests that ants arose in the mid-Cretaceous period about 110 to 130 million years ago. After the rise of flowering plants about 100 million years ago they diversified and assumed ecological dominance around 60 million years ago. In 1966, E. O. Wilson and his colleagues identified the fossil remains of an ant ("Sphecomyrma freyi") that lived in the Cretaceous period. The specimen, trapped in amber dating back to more than 80 million years ago, has features of both ants and wasps. "Sphecomyrma" was probably a ground forager but some suggest on the basis of groups such as the Leptanillinae and Martialinae that primitive ants were likely to have been predators under the soil surface. During the Cretaceous period, only a few species of primitive ants ranged widely on the Laurasian super-continent (the northern hemisphere). They were scarce in comparison to other insects, representing about 1% of the insect population. Ants became dominant after adaptive radiation at the beginning of the Tertiary period. By the Oligocene and Miocene ants had come to represent 20–40% of all insects found in major fossil deposits. Of the species that lived in the Eocene epoch, approximately one in ten genera survive to the present. Genera surviving today comprise 56% of the genera in Baltic amber fossils (early Oligocene), and 92% of the genera in Dominican amber fossils (apparently early Miocene). Termites, though sometimes called "white ants", are not ants and belong to the order Isoptera. The termites are actually more closely related to cockroaches and mantids. The fact that ants and termites are both eusocial came about by Convergent evolution. Velvet ants look like large ants, but are wingless female wasps. Etymology. The word "ant" is derived from "ante" of Middle English which is derived from "æmette" and "emmett" of Old English and is related to the Old High German "āmeiza" from which comes "Ameise", the German word for ant. The family name "Formicidae" is derived from the Latin "formīca" ("ant") from which derived Portuguese "formiga", Spanish "hormiga", Romanian "furnică", French "fourmi", etc. Distribution and diversity. Ants are found on all continents except Antarctica and only a few large islands such as Greenland, Iceland, parts of Polynesia and the Hawaiian Islands lack native ant species. Ants occupy a wide range of ecological niches, and are able to exploit a wide range of food resources either as direct or indirect herbivores, predators and scavengers. Most species are omnivorous generalists but a few are specialist feeders. Their ecological dominance may be measured by their biomass, and estimates in different environments suggest that they contribute 15–20% (on average and nearly 25% in the tropics) of the total terrestrial animal biomass, which exceeds that of the vertebrates. Ants range in size from. Their colours vary; most are red or black, green is less common, and some tropical species have a metallic lustre. More than 12,000 species are currently known (with upper estimates of about 14,000), with the greatest diversity in the tropics. Taxonomic studies continue to resolve the classification and systematics of ants. Online databases of ant species, including AntBase and the Hymenoptera Name Server, help to keep track of the known and newly described species. The relative ease with which ants can be sampled and studied in ecosystems has made them useful as indicator species in biodiversity studies. Morphology. Ants are distinct in their morphology from other insects in having elbowed antennae, metapleural glands, and a strong constriction of their second abdominal segment into a node-like petiole. The head, mesosoma and metasoma or gaster are the three distinct body segments. The petiole forms a narrow waist between their mesosoma (thorax plus the first abdominal segment, which is fused to it) and gaster (abdomen less the abdominal segments in the petiole). The petiole can be formed by one or two nodes (the second alone, or the second and third abdominal segments). Like other insects, ants have an exoskeleton, an external covering that provides a protective casing around the body and a point of attachment for muscles, in contrast to the internal skeletons of humans and other vertebrates. Insects do not have lungs; oxygen and other gases like carbon dioxide pass through their exoskeleton through tiny valves called spiracles. Insects also lack closed blood vessels; instead, they have a long, thin, perforated tube along the top of the body (called the "dorsal aorta") that functions like a heart, and pumps haemolymph towards the head, thus driving the circulation of the internal fluids. The nervous system consists of a ventral nerve cord that runs the length of the body, with several ganglia and branches along the way reaching into the extremities of the appendages. An ant's head contains many sensory organs. Like most insects, ants have compound eyes made from numerous tiny lenses attached together. Ants' eyes are good for acute movement detection but do not give a high resolution. They also have three small ocelli (simple eyes) on the top of the head that detect light levels and polarisation. Compared to vertebrates, most ants have poor-to-mediocre eyesight and a few subterranean species are completely blind. Some ants such as Australia's bulldog ant, however, have exceptional vision. Two antennae ("feelers") are attached to the head; these organs detect chemicals, air currents and vibrations; they are also used to transmit and receive signals through touch. The head has two strong jaws, the mandibles, used to carry food, manipulate objects, construct nests, and for defence. In some species a small pocket (infrabuccal chamber) inside the mouth stores food, so it can be passed to other ants or their larvae. All six legs are attached to the mesosoma ("thorax"). A hooked claw at the end of each leg helps ants to climb and hang onto surfaces. Most queens and male ants have wings; queens shed the wings after the nuptial flight, leaving visible stubs, a distinguishing feature of queens. However, wingless queens (ergatoids) and males occur in a few species. The metasoma (the "abdomen") of the ant houses important internal organs, including those of the reproductive, respiratory (tracheae) and excretory systems. Workers of many species have their egg-laying structures modified into stings that are used for subduing prey and defending their nests. Polymorphism. In the colonies of a few ant species, there are physical castes—workers in distinct size-classes, called minor, median, and major workers. Often the larger ants have disproportionately larger heads, and correspondingly stronger mandibles. Such individuals are sometimes called "soldier" ants because their stronger mandibles make them more effective in fighting, although they are still workers and their "duties" typically do not vary greatly from the minor or median workers. In a few species the median workers are absent, creating a sharp divide between the minors and majors. Weaver ants, for example, have a distinct bimodal size distribution. Some other species show continuous variation in the size of workers. The smallest and largest workers in "Pheidologeton diversus" show nearly a 500-fold difference in their dry-weights. Workers cannot mate; however, because of the haplodiploid sex-determination system in ants, workers of a number of species can lay unfertilised eggs that become fully fertile haploid males. The role of workers may change with their age and in some species, such as honeypot ants, young workers are fed until their gasters are distended, and act as living food storage vessels. These food storage workers are called "repletes". This polymorphism in morphology and behaviour of workers was initially thought to be determined by environmental factors such as nutrition and hormones which led to different developmental paths; however, genetic differences between worker castes have been noted in "Acromyrmex" sp. These polymorphisms are caused by relatively small genetic changes; differences in a single gene of "Solenopsis invicta" can decide whether the colony will have single or multiple queens. The Australian jack jumper ant ("Myrmecia pilosula"), has only a single pair of chromosomes (males have just one chromosome as they are haploid), the lowest number known for any animal, making it an interesting subject for studies in the genetics and developmental biology of social insects. Development and reproduction. The life of an ant starts from an egg. If the egg is fertilised, the progeny will be female (diploid); if not, it will be male (haploid). Ants develop by complete metamorphosis with the larval stages passing through a pupal stage before emerging as an adult. The larva is largely immobile and is fed and cared for by workers. Food is given to the larvae by trophallaxis, a process in which an ant regurgitates liquid food held in its crop. This is also how adults share food, stored in the "social stomach", among themselves. Larvae may also be provided with solid food such as trophic eggs, pieces of prey and seeds brought back by foraging workers and may even be transported directly to captured prey in some species. The larvae grow through a series of moults and enter the pupal stage. The pupa has the appendages free and not fused to the body as in a butterfly pupa. The differentiation into queens and workers (which are both female), and different castes of workers (when they exist), is determined by the nutrition the larvae obtain. Larvae and pupae need to be kept at fairly constant temperatures to ensure proper development, and so are often moved around the various brood chambers within the colony. A new worker spends the first few days of its adult life caring for the queen and young. It then graduates to digging and other nest work, and later to defending the nest and foraging. These changes are sometimes fairly sudden, and define what are called temporal castes. An explanation for the sequence is suggested by the high casualties involved in foraging, making it an acceptable risk only for ants that are older and are likely to die soon of natural causes. Most ant species have a system in which only the queen and breeding females have the ability to mate. Contrary to popular belief, some ant nests have multiple queens while others can exist without queens. Workers with the ability to reproduce are called "gamergates" and colonies that lack queens are then called gamergate colonies; colonies with queens are said to be queen-right. The winged male ants, called drones, emerge from pupae along with the breeding females (although some species, like army ants, have wingless queens), and do nothing in life except eat and mate. During the short breeding period, the reproductives, excluding the colony queen, are carried outside where other colonies of similar species are doing the same. Then, all the winged breeding ants take flight. Mating occurs in flight and the males die shortly afterwards. Females of some species mate with multiple males. Mated females then seek a suitable place to begin a colony. There, they break off their wings and begin to lay and care for eggs. The females store the sperm they obtain during their nuptial flight to selectively fertilise future eggs. The first workers to hatch are weak and smaller than later workers, but they begin to serve the colony immediately. They enlarge the nest, forage for food and care for the other eggs. This is how new colonies start in most species. Species that have multiple queens may have a queen leaving the nest along with some workers to found a colony at a new site, a process akin to swarming in honeybees. Ant colonies can be long-lived. The queens can live for up to 30 years, and workers live from 1 to 3 years. Males, however, are more transitory, and survive only a few weeks. Ant queens are estimated to live 100 times longer than solitary insects of a similar size. Ants are active all year long in the tropics but, in cooler regions, survive the winter in a state of dormancy or inactivity. The forms of inactivity are varied and some temperate species have larvae going into the inactive state (diapause), while in others, the adults alone pass the winter in a state of reduced activity. Communication. Ants communicate with each other using pheromones. These chemical signals are more developed in ants than in other hymenopteran groups. Like other insects, ants perceive smells with their long, thin and mobile antennae. The paired antennae provide information about the direction and intensity of scents. Since most ants live on the ground, they use the soil surface to leave pheromone trails that can be followed by other ants. In species that forage in groups, a forager that finds food marks a trail on the way back to the colony; this trail is followed by other ants, these ants then reinforce the trail when they head back with food to the colony. When the food source is exhausted, no new trails are marked by returning ants and the scent slowly dissipates. This behaviour helps ants deal with changes in their environment. For instance, when an established path to a food source is blocked by an obstacle, the foragers leave the path to explore new routes. If an ant is successful, it leaves a new trail marking the shortest route on its return. Successful trails are followed by more ants, reinforcing better routes and gradually finding the best path. Ants use pheromones for more than just making trails. A crushed ant emits an alarm pheromone that sends nearby ants into an attack frenzy and attracts more ants from further away. Several ant species even use "propaganda pheromones" to confuse enemy ants and make them fight among themselves. Pheromones are produced by a wide range of structures including Dufour's glands, poison glands and glands on the hindgut, pygidium, rectum, sternum and hind tibia. Pheromones are also exchanged mixed with food and passed by trophallaxis, transferring information within the colony. This allows other ants to detect what task group ("e.g.", foraging or nest maintenance) other colony members belong to. In ant species with queen castes, workers begin to raise new queens in the colony when the dominant queen stops producing a specific pheromone. Some ants produce sounds by stridulation, using the gaster segments and their mandibles. Sounds may be used to communicate with colony members or with other species. Defence===. Ants attack and defend themselves by biting and, in many species, by stinging, often injecting or spraying chemicals like formic acid. Bullet ants ("Paraponera"), located in Central and South America, are considered to have the most painful sting of any insect, although it is usually not fatal to humans. This sting is given the highest rating on the Schmidt Sting Pain Index. The sting of Jack jumper ants can be fatal, and an antivenin has been developed. Fire ants, "Solenopsis" spp., are unique in having a poison sac containing piperidine alkaloids. Their stings are painful and can be dangerous to hypersensitive people. Trap-jaw ants of the genus "Odontomachus" are equipped with mandibles called trap-jaws, which snap shut faster than any other predatory appendages within the animal kingdom. One study of "Odontomachus bauri" recorded peak speeds of between 126 and 230 h (78 143 mph), with the jaws closing within 130 microseconds on average. The ants were also observed to use their jaws as a catapult to eject intruders or fling themselves backwards to escape a threat. Before the strike, the ant opens its mandibles extremely widely and locks them in this position by an internal mechanism. Energy is stored in a thick band of muscle and explosively released when triggered by the stimulation of sensory hairs on the inside of the mandibles. The mandibles also permit slow and fine movements for other tasks. Trap-jaws are also seen in the following genera: "Anochetus", "Orectognathus", and "Strumigenys", plus some members of the Dacetini tribe, which are viewed as examples of convergent evolution. In addition to defence against predators, ants need to protect their colonies from pathogens. Some worker ants maintain the hygiene of the colony and their activities include undertaking or "necrophory", the disposal of dead nest-mates. Oleic acid has been identified as the compound released by dead ants that triggers undertaking behaviour in "Atta mexicana". Nests may be protected from physical threats such as flooding and over-heating by elaborate nest architecture. Workers of "Cataulacus muticus", an arboreal species that lives in plant hollows, respond to flooding by drinking water inside the nest, and excreting it outside. Learning. Many animals can learn behaviours by imitation but ants may be the only group apart from mammals where interactive teaching has been observed. A knowledgeable forager of "Temnothorax albipennis" leads a naive nest-mate to newly discovered food by the excruciatingly slow process of tandem running. The follower obtains knowledge through its leading tutor. Both leader and follower are acutely sensitive to the progress of their partner with the leader slowing down when the follower lags, and speeding up when the follower gets too close. Controlled experiments with colonies of "Cerapachys biroi" suggest that individuals may choose nest roles based on their previous experience. An entire generation of identical workers was divided into two groups whose outcome in food foraging was controlled. One group was continually rewarded with prey, while it was made certain that the other failed. As a result, members of the successful group intensified their foraging attempts while the unsuccessful group ventured out less and less. A month later, the successful foragers continued in their role while the others moved to specialise in brood care. Nest construction. Complex nests are built by many ants, but other species are nomadic and do not build permanent structures. Ants may form subterranean nests or build them on trees. These nests can be found in the ground, under stones or logs, inside logs, hollow stems or even acorns. The materials used for construction include soil and plant matter, and ants carefully select their nest sites; "Temnothorax albipennis" will avoid sites with dead ants, as these may indicate the presence of pests or disease. They are quick to abandon established nests at the first sign of threats. The army ants of South America and the driver ants of Africa do not build permanent nests, but instead alternate between nomadism and stages where the workers form a temporary nest (bivouac) from their own bodies, by holding each other together. Weaver ant ("Oecophylla" spp.) workers build nests in trees by attaching leaves together, first pulling them together with bridges of workers and then inducing their larvae to produce silk as they are moved along the leaf edges. Similar forms of nest construction are seen in some species of "Polyrhachis". Food cultivation. Most ants are generalist predators, scavengers and indirect herbivores, but a few have evolved specialised ways of obtaining nutrition. Leafcutter ants ("Atta" and "Acromyrmex") feed exclusively on a fungus that grows only within their colonies. They continually collect leaves which are taken to the colony, cut into tiny pieces and placed in fungal gardens. Workers specialise in tasks according to their sizes. The largest ants cut stalks, smaller workers chew the leaves and the smallest tend the fungus. Leafcutter ants are sensitive enough to recognise the reaction of the fungus to different plant material, apparently detecting chemical signals from the fungus. If a particular type of leaf is toxic to the fungus the colony will no longer collect it. The ants feed on structures produced by the fungi called "gongylidia". Symbiotic bacteria on the exterior surface of the ants produce antibiotics that kill bacteria that may harm the fungi. Navigation. Foraging ants travel distances of up to from their nest and usually find their way back using scent trails. Some ants forage at night. Day foraging ants in hot and arid regions face death by desiccation, so the ability to find the shortest route back to the nest reduces that risk. Diurnal desert ants ("Cataglyphis fortis") use visual landmarks in combination with other cues to navigate. In the absence of visual landmarks, the closely related Sahara desert ant ("Cataglyphis bicolor") navigates by keeping track of direction as well as distance travelled, like an internal pedometer that counts how many steps they take in each direction. They integrate this information to find the shortest route back to their nest. Several species of ants are able to use the Earth's magnetic field. Ants' compound eyes have specialised cells that detect polarised light from the Sun, which is used to determine direction. Locomotion. Worker ants do not have wings and reproductive females lose their wings after their mating flights in order to begin their colonies. Therefore, unlike their wasp ancestors, most ants travel by walking. Some species are capable of leaping. For example, Jerdon's jumping ant ("Harpegnathos saltator") is able to jump by synchronising the action of its mid and hind pairs of legs. There are several species of gliding ant including "Cephalotes atratus"; this may be a common trait among most arboreal ants. Ants with this ability are able to control the direction of their descent while falling. Other species of ants can form chains to bridge gaps over water, underground, or through spaces in vegetation. Some species also form floating rafts that help them survive floods. These rafts may also have a role in allowing ants to colonise islands. "Polyrhachis sokolova", a species of ant found in Australian mangrove swamps, can swim and live in underwater nests. Since they lack gills, they breathe in trapped pockets of air in the submerged nests. Cooperation and competition. Not all ants have the same kind of societies. The Australian bulldog ants are among the biggest and most basal (primitive) of ants. Like virtually all ants they are eusocial, but their social behaviour is poorly developed compared to other species. Each individual hunts alone, using its large eyes instead of its chemical senses to find prey. Some species (such as "Tetramorium caespitum") attack and take over neighbouring ant colonies. Others are less expansionist but just as aggressive; they invade colonies to steal eggs or larvae, which they either eat or raise as workers slaves. Extreme specialists among these slave-raiding ants, such as the Amazon ants, are incapable of feeding themselves and need captured workers to survive. Ants identify kin and nestmates through their scent, which comes from hydrocarbon-laced secretions that coat their exoskeletons. If an ant is separated from its original colony, it will eventually lose the colony scent. Any ant that enters a colony without a matching scent will be attacked. Parasitic ant species enter the colonies of host ants and establish themselves as social parasites; species like "Strumigenys xenos" are entirely parasitic and do not have workers, but instead rely on the food gathered by their "Strumigenys perplexa" hosts. This form of parasitism is seen across many ant genera, but the parasitic ant is usually a species that is closely related to its host. A variety of methods are employed to enter the nest of the host ant. A parasitic queen can enter the host nest before the first brood has hatched, establishing herself prior to development of a colony scent. Other species use pheromones to confuse the host ants or to trick them into carrying the parasitic queen into the nest. Some simply fight their way into the nest. A conflict between the sexes of a species is seen in some species of ants with the reproductives apparently competing to produce offspring that are as closely related to them as possible. The most extreme form involves the production of clonal offspring. An extreme of sexual conflict is seen in "Wasmannia auropunctata", where the queens produce diploid daughters by thelytokous parthenogenesis and males produce clones by a process where a diploid egg loses its maternal contribution to produce haploid males that are clones of the father. Relationships with other organisms. The spider "Myrmarachne plataleoides" (here a female) mimics weaver ants to avoid predators. Ants form symbiotic associations with a range of species, including other ant species, other insects, plants, and fungi. They are preyed on by many animals and even certain fungi. Some arthropod species spend part of their lives within ant nests, either preying on ants, their larvae and eggs, consuming the ants' food stores, or avoiding predators. These inquilines can bear a close resemblance to ants. The nature of this ant mimicry (myrmecomorphy) varies, with some cases involving Batesian mimicry, where the mimic reduces the risk of predation. Others show Wasmannian mimicry, a form of mimicry seen only in inquilines. Aphids and other hemipteran insects secrete a sweet liquid called honeydew when they feed on plant sap. The sugars in honeydew are a high-energy food source, which many ant species collect. In some cases the aphids secrete the honeydew in response to the ants' tapping them with their antennae. The ants in turn keep predators away and will move the aphids between feeding locations. On migrating to a new area, many colonies will take the aphids with them, to ensure a continued supply of honeydew. Ants also tend mealybugs to harvest their honeydew. Mealybugs can become a serious pest of pineapples if ants are present to protect mealybugs from their natural enemies. Myrmecophilous (ant-loving) caterpillars of the family Lycaenidae (e.g., blues, coppers, or hairstreaks) are herded by the ants, led to feeding areas in the daytime, and brought inside the ants' nest at night. The caterpillars have a gland which secretes honeydew when the ants massage them. Some caterpillars produce vibrations and sounds that are perceived by the ants. Other caterpillars have evolved from ant-loving to ant-eating: these myrmecophagous caterpillars secrete a pheromone that makes the ants act as if the caterpillar is one of their own larvae. The caterpillar is then taken into the ants' nest where it feeds on the ant larvae. Fungus-growing ants that make up the tribe Attini, including leafcutter ants, cultivate certain species of fungus in the "Leucoagaricus" or "Leucocoprinus" genera of the Agaricaceae family. In this ant-fungus mutualism, both species depend on each other for survival. The ant "Allomerus decemarticulatus" has evolved a three-way association with the host plant "Hirtella physophora" (Chrysobalanaceae), and a sticky fungus which is used to trap their insect prey. Lemon ants make devil's gardens by killing surrounding plants with their stings and leaving a pure patch of lemon ant	The bicycle'", "'bike'", or "'cycle'" is a pedal-driven, human-powered vehicle with two wheels attached to a frame, one behind the other. Bicycles were introduced in the 19th century and now number about one billion worldwide. They are the principal means of transportation in many regions. They also provide a popular form of recreation, and have been adapted for such uses as children's toys, adult fitness, military and police applications, courier services, and competitive sports. The basic shape and configuration of a typical bicycle has changed little since the first chain-driven model was developed around 1885. Many details have been improved, especially since the advent of modern materials and computer-aided design. These have allowed for a proliferation of specialized designs for particular types of cycling. The bicycle has had a considerable effect on human society, in both the cultural and industrial realms. In its early years, bicycle construction drew on pre-existing technologies; more recently, bicycle technology has, in turn, contributed both to old and new areas. History. Multiple innovators contributed to the history of the bicycle by developing precursor human-powered vehicles. The documented ancestors of today's modern bicycle were known as push bikes (still called push bikes outside of North America), draisines, or hobby horses. Being the first human means of transport to make use of the two-wheeler principle, the draisine (or "mistmashine", "running machine"), invented by the German Baron Karl von Drais, is regarded as the archetype of the bicycle. It was introduced by Drais to the public in Mannheim in summer 1817 and in Paris in 1818. Its rider sat astride a wooden frame supported by two in-line wheels and pushed the vehicle along with his her feet while steering the front wheel. In the early 1860s, Frenchmen Pierre Michaux and Pierre Lallement took bicycle design in a new direction by adding a mechanical crank drive with pedals on an enlarged front wheel. Another French inventor by the name of Douglas Grasso had a failed prototype of Pierre Lallement's bicycle several years earlier. Several why-not-the-rear-wheel inventions followed, the best known being the rod-driven velocipede by Scotsman Thomas McCall in 1869. The French creation, made of iron and wood, developed into the "penny-farthing" (more formally an "ordinary bicycle", a retronym, since there were then no other kind). It featured a tubular steel frame on which were mounted wire spoked wheels with solid rubber tires. These bicycles were difficult to ride due to their very high seat and poor weight distribution. The "dwarf ordinary" addressed some of these faults by reducing the front wheel diameter and setting the seat further back. This necessitated the addition of gearing, effected in a variety of ways, to attain sufficient speed. Having to both pedal and steer via the front wheel remained a problem. J. K. Starley, J. H. Lawson, and Shergold solved this problem by introducing the chain drive (originated by Henry Lawson's unsuccessful "bicyclette"), connecting the frame-mounted pedals to the rear wheel. These models were known as "dwarf safeties", or "safety bicycles", for their lower seat height and better weight distribution. Starley's 1885 Rover is usually described as the first recognizably modern bicycle. Soon, the "seat tube" was added, creating the double-triangle "diamond frame" of the modern bike. Further innovations increased comfort and ushered in a second bicycle craze, the 1890s' "Golden Age of Bicycles". In 1888, Scotsman John Boyd Dunlop introduced the pneumatic tire, which soon became universal. Soon after, the rear freewheel was developed, enabling the rider to coast. This refinement led to the 1898 invention of coaster brakes. Derailleur gears and hand-operated cable-pull brakes were also developed during these years, but were only slowly adopted by casual riders. By the turn of the century, cycling clubs flourished on both sides of the Atlantic, and touring and racing became widely popular. Bicycles and horse buggies were the two mainstays of private transportation just prior to the automobile, and the grading of smooth roads in the late 19th century was stimulated by the widespread advertising, production, and use of these devices. Uses. Bicycles have been and are employed for many uses: Technical aspects. The bicycle has undergone continual adaptation and improvement since its inception. These innovations have continued with the advent of modern materials and computer-aided design, allowing for a proliferation of specialized bicycle types. Types. Bicycles can be categorized in different ways: e.g. by function, by number of riders, by general construction, by gearing or by means of propulsion. The more common types include utility bicycles, mountain bicycles, racing bicycles, touring bicycles, hybrid bicycles, cruiser bicycles, and BMX bicycles. Less common are tandems, lowriders, tall bikes, fixed gear (fixed-wheel), folding models and recumbents (one of which was used to set the IHPVA Hour record). Unicycles, tricycles and quadracycles are not strictly bicycles, as they have respectively one, three and four wheels, but are often referred to informally as "bikes". Dynamics. A bicycle stays upright while moving forward by being steered so as to keep its center of gravity over the wheels. This steering is usually provided by the rider, but under certain conditions may be provided by the bicycle itself. The combined center of mass of a bicycle and its rider must lean into a turn in order successfully navigate it. This lean is induced by a method known as countersteering, which can be performed by the rider turning the handlebars directly with the hands or indirectly by leaning the bicycle. Short-wheelbase or tall bicycles, when braking, can generate enough stopping force at the front wheel in order to flip longitudinally. The act of purposefully using this force to lift the rear wheel and balance on the front without tipping over is a trick known as a stoppie, endo or front wheelie. Performance. The bicycle is extraordinarily efficient in both biological and mechanical terms. The bicycle is the most efficient self-powered means of transportation in terms of energy a person must expend to travel a given distance. From a mechanical viewpoint, up to 99% of the energy delivered by the rider into the pedals is transmitted to the wheels, although the use of gearing mechanisms may reduce this by 10-15%. In terms of the ratio of cargo weight a bicycle can carry to total weight, it is also a most efficient means of cargo transportation. A human traveling on a bicycle at low to medium speeds of around 10-15 mph (15-25 km h), uses only the energy required to walk, is the most energy-efficient means of transport generally available. Air drag, which is proportional to the square of speed, requires dramatically higher power outputs as speeds increase. If the rider is sitting upright, the rider's body creates about 75% of the total drag of the bicycle rider combination. Drag can be reduced by seating the rider in a supine position or a prone position, thus creating a recumbent bicycle or human powered vehicle. Drag can also be reduced by covering the bicycle with an aerodynamic fairing. In addition, the carbon dioxide generated in the production and transportation of the food required by the bicyclist, per mile traveled, is less than 1 10th that generated by energy efficient cars. Construction and parts. In its early years, bicycle construction drew on pre-existing technologies. More recently, bicycle technology has in turn contributed ideas in both old and new areas. Frame. The great majority of today's bicycles have a frame with upright seating which looks much like the first chain-driven bike. Such upright bicycles almost always feature the "diamond frame", a truss consisting of two triangles: the front triangle and the rear triangle. The front triangle consists of the head tube, top tube, down tube and seat tube. The head tube contains the headset, the set of bearings that allows the fork to turn smoothly for steering and balance. The top tube connects the head tube to the seat tube at the top, and the down tube connects the head tube to the bottom bracket. The rear triangle consists of the seat tube and paired chain stays and seat stays. The chain stays run parallel to the chain, connecting the bottom bracket to the rear dropouts. The seat stays connect the top of the seat tube (at or near the same point as the top tube) to the rear dropouts. Historically, women's bicycle frames had a top tube that connected in the middle of the seat tube instead of the top, resulting in a lower standover height at the expense of compromised structural integrity, since this places a strong bending load in the seat tube, and bicycle frame members are typically weak in bending. This design, referred to as a "step-through frame", allows the rider to mount and dismount in a dignified way while wearing a skirt or dress. While some women's bicycles continue to use this frame style, there is also a variation, the "mixte", which splits the top tube into two small top tubes that bypass the seat tube and connect to the rear dropouts. The ease of stepping through is also appreciated by those with limited flexibility or other joint problems. Because of its persistent image as a "women's" bicycle, step-through frames are not common for larger frames. Another style is the recumbent bicycle. These are inherently more aerodynamic than upright versions, as the rider may lean back onto a support and operate pedals that are on about the same level as the seat. The world's fastest bicycle is a recumbent bicycle but this type was banned from competition in 1934 by the Union Cycliste Internationale. Historically, materials used in bicycles have followed a similar pattern as in aircraft, the goal being high strength and low weight. Since the late 1930s alloy steels have been used for frame and fork tubes in higher quality machines. Celluloid found application in mudguards, and aluminum alloys are increasingly used in components such as handlebars, seat post, and brake levers. In the 1980s aluminum alloy frames became popular, and their affordability now makes them common. More expensive carbon fiber and titanium frames are now also available, as well as advanced steel alloys and even bamboo. Drivetrain and gearing. Since cyclists' legs are most efficient over a narrow range of pedaling speeds (cadence), a variable gear ratio helps a cyclist to maintain an optimum pedalling speed while covering varied terrain. As a first approximation, utility bicycles often use a hub gear with a small number (3 to 5) of widely-spaced gears, road bicycles and racing bicycles use derailleur gears with a moderate number (10 to 22) of closely-spaced gears, while mountain bicycles, hybrid bicycles, and touring bicycles use dérailleur gears with a larger number (15 to 30) of moderately-spaced gears, often including an extremely low gear (granny gear) for climbing steep hills. Different gears and ranges of gears are appropriate for different people and styles of cycling. Multi-speed bicycles allow gear selection to suit the circumstances, e.g. it may be comfortable to use a high gear when cycling downhill, a medium gear when cycling on a flat road, and a low gear when cycling uphill. In a lower gear every turn of the pedals leads to fewer rotations of the rear wheel. This allows the energy required to move the same distance to be distributed over more pedal turns, reducing fatigue when riding uphill, with a heavy load, or against strong winds. A higher gear allows a cyclist to make fewer pedal cycles to maintain a given speed, but with more effort per turn of the pedals. The "drivetrain" begins with pedals which rotate the cranks, which are held in axis by the bottom bracket. Most bicycles use a chain to transmit power to the rear wheel. A relatively small number of bicycles use a shaft drive to transmit power. A very small number of bicycles (mainly single-speed bicycles intended for short-distance commuting) use a belt drive as an oil-free way of transmitting power. With a "chain drive" transmission, a "chainring" attached to a crank drives the chain, which in turn rotates the rear wheel via the rear sprocket(s) (cassette or freewheel). There are four gearing options: two-speed hub gear integrated with chain ring, up to 3 chain rings, up to 11 sprockets, hub gear built in to rear wheel (3-speed to 14-speed). The most common options are either a rear hub or multiple chain rings combined with multiple sprockets (other combinations of options are possible but less common). With a "shaft drive" transmission, a gear set at the bottom bracket turns the shaft, which then turns the rear wheel via a gear set connected to the wheel's hub. There is some small loss of efficiency due to the two gear sets needed. The only gearing option with a shaft drive is to use a hub gear. Steering and seating. The handlebars turn the fork and the front wheel via the stem, which rotates within the headset. Three styles of handlebar are common. "Upright handlebars", the norm in Europe and elsewhere until the 1970s, curve gently back toward the rider, offering a natural grip and comfortable upright position. "Drop handlebars" "drop" as they curve forward and down, offering the cyclist best braking power from a more aerodynamic "crouched" position, as well as more upright positions in which the hands grip the brake lever mounts, the forward curves, or the upper flat sections for increasingly upright postures. Mountain bikes generally feature a 'straight handlebar' or 'riser bar' with varying degrees of sweep backwards and centimeters rise upwards, as well as wider widths which can provide better handling due to increased leverage against the wheel. Saddles also vary with rider preference, from the cushioned ones favored by short-distance riders to narrower saddles which allow more room for leg swings. Comfort depends on riding position. With comfort bikes and hybrids the cyclist sits high over the seat, their weight directed down onto the saddle, such that a wider and more cushioned saddle is preferable. For racing bikes where the rider is bent over, weight is more evenly distributed between the handlebars and saddle, the hips are flexed, and a narrower and harder saddle is more efficient. Differing saddle designs exist for male and female cyclists, accommodating the genders' differing anatomies, although bikes typically are sold with saddles most appropriate for men. A recumbent bicycle has a reclined chair-like seat that some riders find more comfortable than a saddle, especially riders who suffer from certain types of seat, back, neck, shoulder, or wrist pain. Recumbent bicycles may have either under-seat or over-seat steering. Brakes. Modern bicycle "brakes" may be "rim brakes", in which friction pads are compressed against the wheel rims, "internal hub brakes", in which the friction pads are contained within the wheel hubs, "disc brakes", with a separate rotor for braking. Disc brakes are more common on off-road bicycles, tandems and recumbent bicycles than on road-specific bicycles. With hand-operated brakes, force is applied to brake levers mounted on the handlebars and transmitted via Bowden cables or hydraulic lines to the friction pads. A rear hub brake may be either hand-operated or pedal-actuated, as in the back pedal "coaster brakes" which were popular in North America until the 1960s, and are still common in children's bicycles. Track bicycles do not have dedicated brakes. Brakes are not required for riding on a track because all riders ride in the same direction around a track which does not necessitate sharp deceleration. Track riders are still able to slow down because all track bicycles are fixed-gear, meaning that there is no freewheel. Without a freewheel, coasting is impossible, so when the rear wheel is moving, the crank is moving. To slow down, the rider applies resistance to the pedals – this acts as a braking system which can be as effective as a friction-based rear wheel brake, but not as effective as a front wheel brake. Suspension. Bicycle suspension refers to the system or systems used to "suspend" the rider and all or part of the bicycle. This serves two purposes: Bicycle suspensions are used primarily on mountain bicycles, but are also common on hybrid bicycles, and can even be found on some road bicycles, as they can help deal with problematic vibration. Suspension is especially important on recumbent bicycles, since while an upright bicycle rider can stand on the pedals to achieve some of the benefits of suspension, a recumbent rider cannot. Wheels. The wheel axle fits into dropouts in the frame and forks. A pair of wheels may be called a wheelset, especially in the context of ready-built "off the shelf", performance-oriented wheels. Tires vary enormously. Skinny, road-racing tires may be completely smooth, or (slick). On the opposite extreme, off-road tires are much wider and thicker, and usually have a deep tread for gripping in muddy conditions. Accessories, repairs, and tools. Some components, which are often optional accessories on sports bicycles, are standard features on utility bicycles to enhance their usefulness and comfort. Mudguards, or fenders, protect the cyclist and moving parts from spray when riding through wet areas and chainguards protect clothes from oil on the chain while preventing clothing from being caught between the chain and crankset teeth. Kick stands keep a bicycle upright when parked. Front-mounted baskets for carrying goods are often used. Luggage carriers and panniers mounted above the rear tire can be used to carry equipment or cargo. Parents sometimes add rear-mounted child seats and or an auxiliary saddle fitted to the crossbar to transport children. "Toe-clips" and "toestraps" and clipless pedals help keep the foot locked in the proper position on the pedals, and enable the cyclist to pull as well as push the pedals—although not without their hazards, eg. may lock foot in when needed to prevent a fall. Technical accessories include cyclocomputers for measuring speed, distance, etc. Other accessories include lights, reflectors, security locks, mirror, water bottles and cages, and bell. Bicycle helmets may help reduce injury in the event of a collision or accident, and a certified helmet is legally required for some riders in some jurisdictions. Helmets are classified as an accessory or an item of clothing by others. Many cyclists carry "tool kits". These may include a tire patch kit (which, in turn, may contain any combination of a hand pump or CO2 Pump, tire levers, spare tubes, self-adhesive patches, or tube-patching material, an adhesive, a piece of sandpaper or a metal grater (to roughing the tube surface to be patched), and sometimes even a block of French chalk.), wrenches, hex keys, screwdrivers, and a chain tool. There are also cycling specific multi-tools that combine many of these implements into a single compact device. More specialized bicycle components may require more complex tools, including proprietary tools specific for a given manufacturer. Some bicycle parts, particularly hub-based gearing systems, are complex, and many cyclists prefer to leave maintenance and repairs to professional bicycle mechanics. In some areas it is possible to purchase road-side assistance from companies such as the Better World Club. Other cyclists maintain their own bicycles, perhaps as part of their enjoyment of the hobby of cycling or simply for economic reasons. The ability to repair and maintain your own bicycle is also celebrated within the DIY movement. Standards. A number of formal and industry standards exist for bicycle components to help make spare parts exchangeable and to maintain a minimum product safety. The International Organization for Standardization, ISO, has a special technical committee for cycles, TC149, that has the following scope: "Standardization in the field of cycles, their components and accessories with particular reference to terminology, testing methods and requirements for performance and safety, and interchangeability." CEN, European Committee for Standardisation, also has a specific Technical Committee, TC333, that defines European standards for cycles. Their mandate states that EN cycle standards shall harmonize with ISO standards. Some CEN cycle standards were developed before ISO published their standards, leading to strong European influences in this area. European cycle standards tend to describe minimum safety requirements, while ISO standards have historically harmonized parts geometry. Parts. For details on specific bicycle parts, see list of bicycle parts and. Social and historical aspects. The bicycle has had a considerable effect on human society, in both the cultural and industrial realms. In daily life. Around the turn of the 20th century, bicycles reduced crowding in inner-city tenements by allowing workers to commute from more spacious dwellings in the suburbs. They also reduced dependence on horses. Bicycles allowed people to travel for leisure into the country, since bicycles were three times as energy efficient as walking and three to four times as fast. Recently, several European cities have implemented successful schemes known as community bicycle programs or bike-sharing. These initiatives complement a city's public transport system and offer an alternative to motorized traffic to help reduce congestion and pollution. Users take a bicycle at a parking station, use it for a limited amount of time, and then return it to the same or different station. Examples include Bicing in Barcelona, Vélo'v in Lyon and Vélib' in Paris. In cities where the bicycle is not an integral part of the planned transportation system, commuters often use bicycles as elements of a mixed-mode commute, where the bike is used to travel to and from train stations or other forms of rapid transit. Folding bicycles are useful in these scenarios, as they are less cumbersome when carried aboard. Los Angeles removed a small amount of seating on some trains to make more room for bicycles and wheel chairs. Bicycles offer an important mode of transport in many developing countries. Until recently, bicycles have been a staple of everyday life throughout Asian countries. They are the most frequently used method of transport for commuting to work, school, shopping, and life in general. As a result, bicycles there are almost always equipped with baskets. Female emancipation. The diamond-frame safety bicycle gave women unprecedented mobility, contributing to their emancipation in Western nations. As bicycles became safer and cheaper, more women had access to the personal freedom they embodied, and so the bicycle came to symbolize the New Woman of the late 19th century, especially in Britain and the United States. The bicycle was recognized by 19th-century feminists and suffragists as a "freedom machine" for women. American Susan B. Anthony said in a "New York World" interview on February 2 1896: "Let me tell you what I think of bicycling. I think it has done more to emancipate women than anything else in the world. It gives women a feeling of freedom and self-reliance. I stand and rejoice every time I see a woman ride by on a wheel...the picture of free, untrammeled womanhood." In 1895 Frances Willard, the tightly-laced president of the Women’s Christian Temperance Union, wrote a book called "How I Learned to Ride the Bicycle", in which she praised the bicycle she learned to ride late in life, and which she named "Gladys", for its "gladdening effect" on her health and political optimism. Willard used a cycling metaphor to urge other suffragists to action, proclaiming, "I would not waste my life in friction when it could be turned into momentum." Male anger at the freedom symbolized by the New (bicycling) Woman was demonstrated when the male undergraduates of Cambridge University showed their opposition to the admission of women as full members of the university by hanging a woman bicyclist in effigy in the main town square. This was as late as 1897. The bicycle craze in the 1890s also led to a movement for so-called rational dress, which helped liberate women from corsets and ankle-length skirts and other restrictive garments, substituting the then-shocking bloomers. Economic implications. Bicycle manufacturing proved to be a training ground for other industries and led to the development of advanced metalworking techniques, both for the frames themselves and for special components such as ball bearings, washers, and sprockets. These techniques later enabled skilled metalworkers and mechanics to develop the components used in early automobiles and aircraft. They also served to teach the industrial models later adopted, including mechanization and mass production (later copied and adopted by Ford and General Motors), vertical integration (also later copied and adopted by Ford), aggressive advertising (as much as 10% of all advertising in U.S. periodicals in 1898 was by bicycle makers), lobbying for better roads (which had the side benefit of acting as advertising, and of improving sales by providing more places to ride), all first practised by Pope. In addition, bicycle makers adopted the annual model change (later derided as planned obsolescence, and usually credited to General Motors), which proved very successful. Furthermore, bicycles were an early example of conspicuous consumption, being adopted by the fashionable elites. In addition, by serving as a platform for accessories, which could ultimately cost more than the bicycle itself, it paved the way for the likes of the Barbie doll. Moreover, they helped create, or enhance, new kinds of businesses, such as bicycle messengers, travelling seamstresses, riding academies, and racing rinks (Their board tracks were later adapted to early motorcycle and automobile racing.) Also, there were a variety of new inventions, such as spoke tighteners, and specialized lights, socks and shoes, and even cameras (such as the Eastman Company's "Poco"). Probably the best known and most widely used of these inventions, adopted well beyond cycling, is Charles Bennett's Bike Web, which came to be called the "jock strap". They also presaged a move away from public transit that would explode with the introduction of the automobile. This liberation would be repeated again with the appearance of the snowmobile. J. K. Starley's company became the Rover Cycle Company Ltd. in the late 1890s, and then simply the Rover Company when it started making cars. The Morris Motor Company (in Oxford) and Škoda also began in the bicycle business, as did the Wright brothers. Alistair Craig, whose company eventually emerged to become the engine manufacturers Ailsa Craig, also started from manufacturing bicycles, in Glasgow in March 1885. In general, U.S. and European cycle manufacturers used to assemble cycles from their own frames and components made by other companies, although very large companies (such as Raleigh) used to make almost every part of a bicycle (including bottom brackets, axles, etc.) In recent years, those bicycle makers have greatly changed their methods of production. Now, almost none of them produce their own frames. Many newer or smaller companies only design and market their products; the actual production is done by Asian companies. For example, some 60% of the world's bicycles are now being made in China. Despite this shift in production, as nations such as China and India become more wealthy, their own use of bicycles has declined due to the increasing affordability of cars and motorcycles. One of the major reasons for the proliferation of Chinese-made bicycles in foreign markets is the lower cost of labor in China. One of the profound economic implications of bicycle use is that it liberates the user from oil consumption (Ballantine, 1972). H.G. Wells said: “Every time I see an adult on a bicycle, I no longer despair for the future of the human race.” (Quotegarden.com). The bicycle is a inexpensive, fast, healthy and environmentally friendly mode of transport (Illich, 1974) Legal requirements. Early in its development, like in the case of automobiles, there were restrictions on the operation of bicycles. Along with advertising, and to gain free publicity, Albert A. Pope litigated on behalf of cyclists The 1968 Vienna Convention on Road Traffic of the United Nations considers a bicycle to be a vehicle, and a person controlling a bicycle (whether actually riding or not) is considered an operator. The traffic codes of many countries reflect these definitions and demand that a bicycle satisfy certain legal requirements, sometimes even including licensing, before it can be used on public roads. In many jurisdictions, it is an offense to use a bicycle that is not in roadworthy condition. In most jurisdictions, bicycles must have functioning front and rear lights when ridden after dark. As some generator or dynamo-driven lamps only operate while moving, rear reflectors are frequently also mandatory. Since a moving bicycle makes little noise, some countries insist that bicycles have a warning bell for use when approaching pedestrians, equestrians, and other cyclists. See also. "'Special uses and related vehicle types'" References. Other authors: Eddie Borysewicz, Greg LeMond, Davis Phinney, Connie Carpenter.