ratio of word probabilities predicted from brain for eye and bicycle

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eye

bicycle

top 10 words in brain distribution (in article):
cell light animal produce water human form muscle body type
top 10 words in brain distribution (in article):
fiber city build town state century Unite produce production country
top 10 words in brain distribution (not in article):
drink lamp wine beer bottle tissue bone structure process valve
top 10 words in brain distribution (not in article):
sheep house wool cotton fabric store street weave hamlet pig
times more probable under eye 30 20 10 6 4 2.5 1.25 1 1.25 2.5 4 6 10 20 30 times more probable under bicycle
(words not in the model)
Eyes'" are organs that detect light, and send signals along the optic nerve to the visual and other areas of the brain. Complex optical systems with resolving power have come in ten fundamentally different forms, and 96% of animal species possess a complex optical system. Image-resolving eyes are present in cnidaria, mollusks, chordates, annelids and arthropods. The simplest "eyes", in even unicellular organisms, do nothing but detect whether the surroundings are light or dark, which is sufficient for the entrainment of circadian rhythms. From more complex eyes, retinal photosensitive ganglion cells send signals along the retinohypothalamic tract to the suprachiasmatic nuclei to effect circadian adjustment. Overview. Complex eyes can distinguish shapes and colors. The visual fields of many organisms, especially predators, involve large areas of binocular vision to improve depth perception; in other organisms, eyes are located so as to maximise the field of view, such as in rabbits and horses. The first proto-eyes evolved among animals 540 million years ago, about the time of the so-called Cambrian explosion. The last common ancestor of animals possessed the biochemical toolkit necessary for vision, and more advanced eyes have evolved in 96% of animal species in 6 of the thirty-something main phyla. In most vertebrates and some mollusks, the eye works by allowing light to enter it and project onto a light-sensitive panel of cells, known as the retina, at the rear of the eye. The cone cells (for color) and the rod cells (for low-light contrasts) in the retina detect and convert light into neural signals for vision. The visual signals are then transmitted to the brain via the optic nerve. Such eyes are typically roughly spherical, filled with a transparent gel-like substance called the vitreous humour, with a focusing lens and often an iris; the relaxing or tightening of the muscles around the iris change the size of the pupil, thereby regulating the amount of light that enters the eye, and reducing aberrations when there is enough light. The eyes of cephalopods, fish, amphibians and snakes usually have fixed lens shapes, and focusing vision is achieved by telescoping the lens similar to how a camera focuses. Compound eyes are found among the arthropods and are composed of many simple facets which, depending on the details of anatomy, may give either a single pixelated image or multiple images, per eye. Each sensor has its own lens and photosensitive cell(s). Some eyes have up to 28,000 such sensors, which are arranged hexagonally, and which can give a full 360-degree field of vision. Compound eyes are very sensitive to motion. Some arthropods, including many Strepsiptera, have compound eyes of only a few facets, each with a retina capable of creating an image, creating vision. With each eye viewing a different thing, a fused image from all the eyes is produced in the brain, providing very different, high-resolution images. Possessing detailed hyperspectral color vision, the Mantis shrimp has been reported to have the world's most complex color vision system. Trilobites, which are now extinct, had unique compound eyes. They used clear calcite crystals to form the lenses of their eyes. In this, they differ from most other arthropods, which have soft eyes. The number of lenses in such an eye varied, however: some trilobites had only one, and some had thousands of lenses in one eye. In contrast to compound eyes, simple eyes are those that have a single lens. For example, jumping spiders have a large pair of simple eyes with a narrow field of view, supported by an array of other, smaller eyes for peripheral vision. Some insect larvae, like caterpillars, have a different type of simple eye (stemmata) which gives a rough image. Some of the simplest eyes, called ocelli, can be found in animals like some of the snails, which cannot actually "see" in the normal sense. They do have photosensitive cells, but no lens and no other means of projecting an image onto these cells. They can distinguish between light and dark, but no more. This enables snails to keep out of direct sunlight. In organisms dwelling near deep-sea vents, compound eyes have been secondarily simplified and adapted to spot the infra-red light produced by the hot vents in this way the bearers can spot hot springs and avoid being boiled alive. Evolution. Visual pigments appear to have a common ancestor and were probably involved in circadian rhythms or reproductive timing in simple organisms. Complex vision, associated with dedicated visual organs, or eyes, evolved many times in different lineages. Types of eye. Nature has produced ten different eye layouts indeed every way of capturing an image has evolved at least once in nature, with the exception of zoom and Fresnel lenses. Eye types can be categorized into "simple eyes", with one concave chamber, and "compound eyes", which comprise a number of individual lenses laid out on a convex surface. Note that "simple" does not imply a reduced level of complexity or acuity. Indeed, any eye type can be adapted for almost any behaviour or environment. The only limitations specific to eye types are that of resolution the physics of compound eyes prevents them from achieving a resolution better than 1°. Also, superposition eyes can achieve greater sensitivity than apposition eyes, so are better suited to dark-dwelling creatures. Eyes also fall into two groups on the basis of their photoreceptor's cellular construction, with the photoreceptor cells either being cilliated (as in the vertebrates) or rhabdomic. These two groups are not monophyletic; the cnidaira also possess cilliated cells, Pit eyes. Pit eyes, also known as stemma, are eye-spots which may be set into a pit to reduce the angles of light that enters and affects the eyespot, to allow the organism to deduce the angle of incoming light. Found in about 85% of phyla, these basic forms were probably the precursors to more advanced types of "simple eye". They are small, comprising up to about 100 cells covering about 100 µm. The directionality can be improved by reducing the size of the aperture, by incorporating a reflective layer behind the receptor cells, or by filling the pit with a refractile material. Pinhole eye. The pinhole eye is an "advanced" form of pit eye incorporating these improvements, most notably a small aperture (which may be adjustable) and deep pit. It is only found in the nautiloids. Without a lens to focus the image, it produces a blurry image, and will blur out a point to the size of the aperture. Consequently, nautiloids can't discriminate between objects with an angular separation of less than 11°. Shrinking the aperture would produce a sharper image, but let in less light. Spherical lensed eye. The resolution of pit eyes can be greatly improved by incorporating a material with a higher refractive index to form a lens, which may greatly reduce the blur radius encountered hence increasing the resolution obtainable. The most basic form, still seen in some gastropods and annelids, consists of a lens of one refractive index. A far sharper image can be obtained using materials with a high refractive index, decreasing to the edges this decreases the focal length and thus allows a sharp image to form on the retina. This also allows a larger aperture for a given sharpness of image, allowing more light to enter the lens; and a flatter lens, reducing spherical aberration. Such an inhomogeneous lens is necessary in order for the focal length to drop from about 4 times the lens radius, to 2.5 radii. Heterogeneous eyes have evolved at least eight times four or more times in gastropods, once in the copepods, once in the annelids and once in the cephalopods. No aquatic organisms possess homogeneous lenses; presumably the evolutionary pressure for a heterogeneous lens is great enough for this stage to be quickly "outgrown". This eye creates an image that is sharp enough that motion of the eye can cause significant blurring. To minimize the effect of eye motion while the animal moves, most such eyes have stabilizing eye muscles. The ocelli of insects bear a simple lens, but their focal point always lies behind the retina; consequently they can never form a sharp image. This capitulates the function of the eye. Ocelli (pit-type eyes of arthropods) blur the image across the whole retina, and are consequently excellent at responding to rapid changes in light intensity across the whole visual field this fast response is further accelerated by the large nerve bundles which rush the information to the brain. Focussing the image would also cause the sun's image to be focussed on a few receptors, with the possibility of damage under the intense light; shielding the receptors would block out some light and thus reduce their sensitivity. This fast response has led to suggestions that the ocelli of insects are used mainly in flight, because they can be used to detect sudden changes in which way is up (because light, especially UV light which is absorbed by vegetation, usually comes from above). Weaknesses. One weakness of this eye construction is that chromatic aberration is still quite high although for organisms without color vision, this is a very minor concern. A weakness of the vertebrate eye is the blind spot which results from a gap in the retina where the optic nerve exits at the back of the eye; the cephalopod eye has no blind spot as the retina is in the opposite orientation. Multiple lenses. Some marine organisms bear more than one lens; for instance the copeopod "Pontella" has three. The outer has a parabolic surface, countering the effects of spherical aberration while allowing a sharp image to be formed. "Copilla'"s eyes have two lenses, which move in and out like a telescope. Such arrangements are rare and poorly understood, but represent an interesting alternative construction. An interesting use of multiple lenses is seen in some hunters such as eagles and jumping spiders, which have a refractive cornea (discussed next): these have a negative lens, enlarging the observed image by up to 50% over the receptor cells, thus increasing their optical resolution. Refractive cornea. In the eyes of most terrestrial vertebrates (along with spiders and some insect larvae) the vitreous fluid has a higher refractive index than the air, relieving the lens of the function of reducing the focal length. This has freed it up for fine adjustments of focus, allowing a very high resolution to be obtained. As with spherical lenses, the problem of spherical aberration caused by the lens can be countered either by using an inhomogeneous lens material, or by flattening the lens. Flattening the lens has a disadvantage: the quality of vision is diminished away from the main line of focus, meaning that animals requiring all-round vision are detrimented. Such animals often display an inhomogeneous lens instead. As mentioned above, a refractive cornea is only useful out of water; in water, there is no difference in refractive index between the vitreous fluid and the surrounding water. Hence creatures which have returned to the water penguins and seals, for example lose their refractive cornea and return to lens-based vision. An alternative solution, borne by some divers, is to have a very strong cornea. Reflector eyes. An alternative to a lens is to line the inside of the eye with mirrors", and reflect the image to focus at a central point. The nature of these eyes means that if one were to peer into the pupil of an eye, one would see the same image that the organism would see, reflected back out. Many small organisms such as rotifers, copeopods and platyhelminths use such organs, but these are too small to produce usable images. Some larger organisms, such as scallops, also use reflector eyes. The scallop "Pecten" has up to 100 millimeter-scale reflector eyes fringing the edge of its shell. It detects moving objects as they pass successive lenses. Compound eyes. A compound eye may consist of thousands of individual photoreception units. The image perceived is a combination of inputs from the numerous ommatidia (individual "eye units"), which are located on a convex surface, thus pointing in slightly different directions. Compared with simple eyes, compound eyes possess a very large view angle, and can detect fast movement and, in some cases, the polarization of light. Because the individual lenses are so small, the effects of diffraction impose a limit on the possible resolution that can be obtained. This can only be countered by increasing lens size and number to see with a resolution comparable to our simple eyes, humans would require compound eyes which would each reach the size of their head. Compound eyes fall into two groups: apposition eyes, which form multiple inverted images, and superposition eyes, which form a single erect image. Compound eyes are common in arthropods, and are also present in annelids and some bivalved molluscs. Compound eyes, in arthropods at least, grow at their margins by the addition of new ommatidia. Apposition eyes. Apposition eyes are the most common form of eye, and are presumably the ancestral form of compound eye. They are found in all arthropod groups, although they may have evolved more than once within this phylum. Some annelids and bivalves also have apposition eyes. They are also possessed by "Limulus", the horseshoe crab, and there are suggestions that other chelicerates developed their simple eyes by reduction from a compound starting point. (Some caterpillars appear to have evolved compound eyes from simple eyes in the opposite fashion.) Apposition eyes work by gathering a number of images, one from each eye, and combining them in the brain, with each eye typically contributing a single point of information. The typical apposition eye has a lens focusing light from one direction on the rhabdom, while light from other directions is absorbed by the dark wall of the ommatidium. In the other kind of apposition eye, found in the Strepsiptera, lenses are not fused to one another, and each forms an entire image; these images are combined in the brain. This is called the schizochroal compound eye or the neural superposition eye. Because images are combined additively, this arrangement allows vision under lower light levels. Superposition eyes. The second type is named the superposition eye. The superposition eye is divided into three types; the refracting, the reflecting and the parabolic superposition eye. The refracting superposition eye has a gap between the lens and the rhabdom, and no side wall. Each lens takes light at an angle to its axis and reflects it to the same angle on the other side. The result is an image at half the radius of the eye, which is where the tips of the rhabdoms are. This kind is used mostly by nocturnal insects. In the parabolic superposition compound eye type, seen in arthropods such as mayflies, the parabolic surfaces of the inside of each facet focus light from a reflector to a sensor array. Long-bodied decapod crustaceans such as shrimp, prawns, crayfish and lobsters are alone in having reflecting superposition eyes, which also has a transparent gap but uses corner mirrors instead of lenses. Parabolic superposition. This eye type functions by refracting light, then using a parabolic mirror to focus the image; it combines features of superposition and apposition eyes. Other. Good fliers like flies or honey bees, or prey-catching insects like praying mantis or dragonflies, have specialized zones of ommatidia organized into a fovea area which gives acute vision. In the acute zone the eye are flattened and the facets larger. The flattening allows more ommatidia to receive light from a spot and therefore higher resolution. There are some exceptions from the types mentioned above. Some insects have a so-called single lens compound eye, a transitional type which is something between a superposition type of the multi-lens compound eye and the single lens eye found in animals with simple eyes. Then there is the mysid shrimp "Dioptromysis paucispinosa". The shrimp has an eye of the refracting superposition type, in the rear behind this in each eye there is a single large facet that is three times in diameter the others in the eye and behind this is an enlarged crystalline cone. This projects an upright image on a specialized retina. The resulting eye is a mixture of a simple eye within a compound eye. Another version is the pseudofaceted eye, as seen in Scutigera. This type of eye consists of a cluster of numerous ocelli on each side of the head, organized in a way that resembles a true compound eye. The body of "Ophiocoma wendtii", a type of brittle star, is covered with ommatidia, turning its whole skin into a compound eye. The same is true of many chitons. Relationship to lifestyle. Eyes are generally adapted to the environment and lifestyle of the organism which bears them. For instance, the distribution of photoreceptors tends to match the area in which the highest acuity is required, with horizon-scanning organisms, such as those that live on the African plains, having a horizontal line of high-density ganglia, while tree-dwelling creatures which require good all-round vision tend to have a symmetrical distribution of ganglia, with acuity decreasing outwards from the centre. Of course, for most eye types, it is impossible to diverge from a spherical form, so only the density of optical receptors can be altered. In organisms with compound eyes, it is the number of ommatidia rather than ganglia that reflects the region of highest data acquisition. Optical superposition eyes are constrained to a spherical shape, but other forms of compound eyes may deform to a shape where more ommatidia are aligned to, say, the horizon, without altering the size or density of individual ommatidia. Eyes of horizon-scanning organisms have stalks so they can be easily aligned to the horizon when this is inclined, for example if the animal is on a slope. An extension of this concept is that the eyes of predators typically have a zone of very acute vision at their centre, to assist in the identification of prey. In deep water organisms, it may not be the centre of the eye that is enlarged. The hyperiid amphipods are deep water animals that feed on organisms above them. Their eyes are almost divided into two, with the upper region thought to be involved in detecting the silhouettes of potential prey or predators against the faint light of the sky above. Accordingly, deeper water hyperiids, where the light against which the silhouettes must be compared is dimmer, have larger "upper-eyes", and may lose the lower portion of their eyes altogether. Depth perception can be enhanced by having eyes which are enlarged in one direction; distorting the eye slightly allows the distance to the object to be estimated with a high degree of accuracy. Acuity is higher among male organisms that mate in mid-air, as they need to be able to spot and assess potential mates against a very large backdrop. On the other hand, the eyes of organisms which operate in low light levels, such as around dawn and dusk or in deep water, tend to be larger to increase the amount of light that can be captured. It is not only the shape of the eye that may be affected by lifestyle. Eyes can be the most visible parts of organisms, and this can act as a pressure on organisms to have more transparent eyes at the cost of function. Eyes may be mounted on stalks to provide better all-round vision, by lifting them above an organism's carapace; this also allows them to track predators or prey without moving the head. Acuity. Visual acuity is often measured in cycles per degree (CPD), which measures an angular resolution, or how much an eye can differentiate one object from another in terms of visual angles. Resolution in CPD can be measured by bar charts of different numbers of white black stripe cycles. For example, if each pattern is 1.75 cm wide and is placed at 1 m distance from the eye, it will subtend an angle of 1 degree, so the number of white black bar pairs on the pattern will be a measure of the cycles per degree of that pattern. The highest such number that the eye can resolve as stripes, or distinguish from a gray block, is then the measurement of visual acuity of the eye. For a human eye with excellent acuity, the maximum theoretical resolution would be 50 CPD (1.2 arcminute per line pair, or a 0.35 mm line pair, at 1 m). A rat can resolve only about 1 to 2 CPD. A horse has higher acuity through most of the visual field of its eyes than a human has, but does not match the high acuity of the human eye's central fovea region. Spherical aberration limits the resolution of a 7 mm pupil to about 3 arcminutes per line pair. At a pupil diameter of 3 mm, the spherical aberration is greatly reduced, resulting in an improved resolution of approximately 1.7 arcminutes per line pair. A resolution of 2 arcminutes per line pair, equivalent to a 1 arcminute gap in an optotype, corresponds to 20 20 (normal vision) in humans. Color. All organisms are restricted to a small range of the electromagnetic spectrum; this varies from creature to creature, but is mainly between 400 and 700 nm. This is a rather small section of the electromagnetic spectrum, probably reflecting the submarine evolution of the organ: water blocks out all but two small windows of the EM spectrum, and there has been no evolutionary pressure among land animals to broaden this range. The most sensitive pigment, rhodopsin, has a peak response at 500 nm. Small changes to the genes coding for this protein can tweak the peak response by a few nm; pigments in the lens can also "filter" incoming light, changing the peak response. Many organisms are unable to discriminate between colors, seeing instead in shades of "grey"; color vision necessitates a range of pigment cells which are primarily sensitive to smaller ranges of the spectrum. In primates, geckos, and other organisms, these take the form of cone cells, from which the more sensitive rod cells evolved. Even if organisms are physically capable of discriminating different colors, this does not necessarily mean that they can perceive the different colors; only with behavioral tests can this be deduced. Most organisms with color vision are able to detect ultraviolet light. This high energy light can be damaging to receptor cells. With a few exceptions (snakes, placental mammals), most organisms avoid these effects by having absorbent oil droplets around their cone cells. The alternative, developed by organisms that had lost these oil droplets in the course of evolution, is to make the lens impervious to UV light this precludes the possibility of any UV light being detected, as it does not even reach the retina. Rods and cones. The retina contains two major types of light-sensitive photoreceptor cells used for vision: the rods and the cones. Rods cannot distinguish colors, but are responsible for low-light black-and-white (scotopic) vision; they work well in dim light as they contain a pigment, visual purple, which is sensitive at low light intensity, but saturates at higher (photopic) intensities. Rods are distributed throughout the retina but there are none at the fovea and none at the blind spot. Rod density is greater in the peripheral retina than in the central retina. Cones are responsible for color vision. They require brighter light to function than rods require. There are three types of cones, maximally sensitive to long-wavelength, medium-wavelength, and short-wavelength light (often referred to as red, green, and blue, respectively, though the sensitivity peaks are not actually at these colors). The color seen is the combined effect of stimuli to, and responses from, these three types of cone cells. Cones are mostly concentrated in and near the fovea. Only a few are present at the sides of the retina. Objects are seen most sharply in focus when their images fall on this spot, as when one looks at an object directly. Cone cells and rods are connected through intermediate cells in the retina to nerve fibers of the optic nerve. When rods and cones are stimulated by light, the nerves send off impulses through these fibers to the brain. Pigment. The pigment molecules used in the eye are various, but can be used to define the evolutionary distance between different groups, and can also be an aid in determining which are closely related although problems of convergence do exist. Opsins are the pigments involved in photoreception. Other pigments, such as melanin, are used to shield the photoreceptor cells from light leaking in from the sides. The opsin protein group evolved long before the last common ancestor of animals, and has continued to diversify since. There are two types of opsin involved in vision; c-opsins, which are associated with ciliary-type photoreceptor cells, and r-opsins, associated with rhabdomeric photoreceptor cells. The eyes of vertebrates usually contain cilliary cells with c-opsins, and (bilaterian) invertebrates have rhabdomeric cells in the eye with r-opsins. However, some "ganglion" cells of vertebrates express r-opsins, suggesting that their ancestors used this pigment in vision, and that remnants survive in the eyes. Likewise, c-opsins have been found to be expressed in the "brain" of some invertebrates. They may have been expressed in ciliary cells of larval eyes, which were subsequently resorbed into the brain on metamorphosis to the adult form. C-opsins are also found in some derived bilaterian-invertebrate eyes, such as the pallial eyes of the bivalve molluscs; however, the lateral eyes (which were presumably the ancestral type for this group, if eyes evolved once there) always use r-opsins. Cnidaria, which are an outgroup to the taxa mentioned above, express c-opsins but r-opsins are yet to be found in this group. Incidentally, the melanin produced in the cnidaria is produced in the same fashion as that in vertebrates, suggesting the common descent of this pigment. The bicycle'", "'bike'", or "'cycle'" is a pedal-driven, human-powered vehicle with two wheels attached to a frame, one behind the other. Bicycles were introduced in the 19th century and now number about one billion worldwide. They are the principal means of transportation in many regions. They also provide a popular form of recreation, and have been adapted for such uses as children's toys, adult fitness, military and police applications, courier services, and competitive sports. The basic shape and configuration of a typical bicycle has changed little since the first chain-driven model was developed around 1885. Many details have been improved, especially since the advent of modern materials and computer-aided design. These have allowed for a proliferation of specialized designs for particular types of cycling. The bicycle has had a considerable effect on human society, in both the cultural and industrial realms. In its early years, bicycle construction drew on pre-existing technologies; more recently, bicycle technology has, in turn, contributed both to old and new areas. History. Multiple innovators contributed to the history of the bicycle by developing precursor human-powered vehicles. The documented ancestors of today's modern bicycle were known as push bikes (still called push bikes outside of North America), draisines, or hobby horses. Being the first human means of transport to make use of the two-wheeler principle, the draisine (or "mistmashine", "running machine"), invented by the German Baron Karl von Drais, is regarded as the archetype of the bicycle. It was introduced by Drais to the public in Mannheim in summer 1817 and in Paris in 1818. Its rider sat astride a wooden frame supported by two in-line wheels and pushed the vehicle along with his her feet while steering the front wheel. In the early 1860s, Frenchmen Pierre Michaux and Pierre Lallement took bicycle design in a new direction by adding a mechanical crank drive with pedals on an enlarged front wheel. Another French inventor by the name of Douglas Grasso had a failed prototype of Pierre Lallement's bicycle several years earlier. Several why-not-the-rear-wheel inventions followed, the best known being the rod-driven velocipede by Scotsman Thomas McCall in 1869. The French creation, made of iron and wood, developed into the "penny-farthing" (more formally an "ordinary bicycle", a retronym, since there were then no other kind). It featured a tubular steel frame on which were mounted wire spoked wheels with solid rubber tires. These bicycles were difficult to ride due to their very high seat and poor weight distribution. The "dwarf ordinary" addressed some of these faults by reducing the front wheel diameter and setting the seat further back. This necessitated the addition of gearing, effected in a variety of ways, to attain sufficient speed. Having to both pedal and steer via the front wheel remained a problem. J. K. Starley, J. H. Lawson, and Shergold solved this problem by introducing the chain drive (originated by Henry Lawson's unsuccessful "bicyclette"), connecting the frame-mounted pedals to the rear wheel. These models were known as "dwarf safeties", or "safety bicycles", for their lower seat height and better weight distribution. Starley's 1885 Rover is usually described as the first recognizably modern bicycle. Soon, the "seat tube" was added, creating the double-triangle "diamond frame" of the modern bike. Further innovations increased comfort and ushered in a second bicycle craze, the 1890s' "Golden Age of Bicycles". In 1888, Scotsman John Boyd Dunlop introduced the pneumatic tire, which soon became universal. Soon after, the rear freewheel was developed, enabling the rider to coast. This refinement led to the 1898 invention of coaster brakes. Derailleur gears and hand-operated cable-pull brakes were also developed during these years, but were only slowly adopted by casual riders. By the turn of the century, cycling clubs flourished on both sides of the Atlantic, and touring and racing became widely popular. Bicycles and horse buggies were the two mainstays of private transportation just prior to the automobile, and the grading of smooth roads in the late 19th century was stimulated by the widespread advertising, production, and use of these devices. Uses. Bicycles have been and are employed for many uses: Technical aspects. The bicycle has undergone continual adaptation and improvement since its inception. These innovations have continued with the advent of modern materials and computer-aided design, allowing for a proliferation of specialized bicycle types. Types. Bicycles can be categorized in different ways: e.g. by function, by number of riders, by general construction, by gearing or by means of propulsion. The more common types include utility bicycles, mountain bicycles, racing bicycles, touring bicycles, hybrid bicycles, cruiser bicycles, and BMX bicycles. Less common are tandems, lowriders, tall bikes, fixed gear (fixed-wheel), folding models and recumbents (one of which was used to set the IHPVA Hour record). Unicycles, tricycles and quadracycles are not strictly bicycles, as they have respectively one, three and four wheels, but are often referred to informally as "bikes". Dynamics. A bicycle stays upright while moving forward by being steered so as to keep its center of gravity over the wheels. This steering is usually provided by the rider, but under certain conditions may be provided by the bicycle itself. The combined center of mass of a bicycle and its rider must lean into a turn in order successfully navigate it. This lean is induced by a method known as countersteering, which can be performed by the rider turning the handlebars directly with the hands or indirectly by leaning the bicycle. Short-wheelbase or tall bicycles, when braking, can generate enough stopping force at the front wheel in order to flip longitudinally. The act of purposefully using this force to lift the rear wheel and balance on the front without tipping over is a trick known as a stoppie, endo or front wheelie. Performance. The bicycle is extraordinarily efficient in both biological and mechanical terms. The bicycle is the most efficient self-powered means of transportation in terms of energy a person must expend to travel a given distance. From a mechanical viewpoint, up to 99% of the energy delivered by the rider into the pedals is transmitted to the wheels, although the use of gearing mechanisms may reduce this by 10-15%. In terms of the ratio of cargo weight a bicycle can carry to total weight, it is also a most efficient means of cargo transportation. A human traveling on a bicycle at low to medium speeds of around 10-15 mph (15-25 km h), uses only the energy required to walk, is the most energy-efficient means of transport generally available. Air drag, which is proportional to the square of speed, requires dramatically higher power outputs as speeds increase. If the rider is sitting upright, the rider's body creates about 75% of the total drag of the bicycle rider combination. Drag can be reduced by seating the rider in a supine position or a prone position, thus creating a recumbent bicycle or human powered vehicle. Drag can also be reduced by covering the bicycle with an aerodynamic fairing. In addition, the carbon dioxide generated in the production and transportation of the food required by the bicyclist, per mile traveled, is less than 1 10th that generated by energy efficient cars. Construction and parts. In its early years, bicycle construction drew on pre-existing technologies. More recently, bicycle technology has in turn contributed ideas in both old and new areas. Frame. The great majority of today's bicycles have a frame with upright seating which looks much like the first chain-driven bike. Such upright bicycles almost always feature the "diamond frame", a truss consisting of two triangles: the front triangle and the rear triangle. The front triangle consists of the head tube, top tube, down tube and seat tube. The head tube contains the headset, the set of bearings that allows the fork to turn smoothly for steering and balance. The top tube connects the head tube to the seat tube at the top, and the down tube connects the head tube to the bottom bracket. The rear triangle consists of the seat tube and paired chain stays and seat stays. The chain stays run parallel to the chain, connecting the bottom bracket to the rear dropouts. The seat stays connect the top of the seat tube (at or near the same point as the top tube) to the rear dropouts. Historically, women's bicycle frames had a top tube that connected in the middle of the seat tube instead of the top, resulting in a lower standover height at the expense of compromised structural integrity, since this places a strong bending load in the seat tube, and bicycle frame members are typically weak in bending. This design, referred to as a "step-through frame", allows the rider to mount and dismount in a dignified way while wearing a skirt or dress. While some women's bicycles continue to use this frame style, there is also a variation, the "mixte", which splits the top tube into two small top tubes that bypass the seat tube and connect to the rear dropouts. The ease of stepping through is also appreciated by those with limited flexibility or other joint problems. Because of its persistent image as a "women's" bicycle, step-through frames are not common for larger frames. Another style is the recumbent bicycle. These are inherently more aerodynamic than upright versions, as the rider may lean back onto a support and operate pedals that are on about the same level as the seat. The world's fastest bicycle is a recumbent bicycle but this type was banned from competition in 1934 by the Union Cycliste Internationale. Historically, materials used in bicycles have followed a similar pattern as in aircraft, the goal being high strength and low weight. Since the late 1930s alloy steels have been used for frame and fork tubes in higher quality machines. Celluloid found application in mudguards, and aluminum alloys are increasingly used in components such as handlebars, seat post, and brake levers. In the 1980s aluminum alloy frames became popular, and their affordability now makes them common. More expensive carbon fiber and titanium frames are now also available, as well as advanced steel alloys and even bamboo. Drivetrain and gearing. Since cyclists' legs are most efficient over a narrow range of pedaling speeds (cadence), a variable gear ratio helps a cyclist to maintain an optimum pedalling speed while covering varied terrain. As a first approximation, utility bicycles often use a hub gear with a small number (3 to 5) of widely-spaced gears, road bicycles and racing bicycles use derailleur gears with a moderate number (10 to 22) of closely-spaced gears, while mountain bicycles, hybrid bicycles, and touring bicycles use dérailleur gears with a larger number (15 to 30) of moderately-spaced gears, often including an extremely low gear (granny gear) for climbing steep hills. Different gears and ranges of gears are appropriate for different people and styles of cycling. Multi-speed bicycles allow gear selection to suit the circumstances, e.g. it may be comfortable to use a high gear when cycling downhill, a medium gear when cycling on a flat road, and a low gear when cycling uphill. In a lower gear every turn of the pedals leads to fewer rotations of the rear wheel. This allows the energy required to move the same distance to be distributed over more pedal turns, reducing fatigue when riding uphill, with a heavy load, or against strong winds. A higher gear allows a cyclist to make fewer pedal cycles to maintain a given speed, but with more effort per turn of the pedals. The "drivetrain" begins with pedals which rotate the cranks, which are held in axis by the bottom bracket. Most bicycles use a chain to transmit power to the rear wheel. A relatively small number of bicycles use a shaft drive to transmit power. A very small number of bicycles (mainly single-speed bicycles intended for short-distance commuting) use a belt drive as an oil-free way of transmitting power. With a "chain drive" transmission, a "chainring" attached to a crank drives the chain, which in turn rotates the rear wheel via the rear sprocket(s) (cassette or freewheel). There are four gearing options: two-speed hub gear integrated with chain ring, up to 3 chain rings, up to 11 sprockets, hub gear built in to rear wheel (3-speed to 14-speed). The most common options are either a rear hub or multiple chain rings combined with multiple sprockets (other combinations of options are possible but less common). With a "shaft drive" transmission, a gear set at the bottom bracket turns the shaft, which then turns the rear wheel via a gear set connected to the wheel's hub. There is some small loss of efficiency due to the two gear sets needed. The only gearing option with a shaft drive is to use a hub gear. Steering and seating. The handlebars turn the fork and the front wheel via the stem, which rotates within the headset. Three styles of handlebar are common. "Upright handlebars", the norm in Europe and elsewhere until the 1970s, curve gently back toward the rider, offering a natural grip and comfortable upright position. "Drop handlebars" "drop" as they curve forward and down, offering the cyclist best braking power from a more aerodynamic "crouched" position, as well as more upright positions in which the hands grip the brake lever mounts, the forward curves, or the upper flat sections for increasingly upright postures. Mountain bikes generally feature a 'straight handlebar' or 'riser bar' with varying degrees of sweep backwards and centimeters rise upwards, as well as wider widths which can provide better handling due to increased leverage against the wheel. Saddles also vary with rider preference, from the cushioned ones favored by short-distance riders to narrower saddles which allow more room for leg swings. Comfort depends on riding position. With comfort bikes and hybrids the cyclist sits high over the seat, their weight directed down onto the saddle, such that a wider and more cushioned saddle is preferable. For racing bikes where the rider is bent over, weight is more evenly distributed between the handlebars and saddle, the hips are flexed, and a narrower and harder saddle is more efficient. Differing saddle designs exist for male and female cyclists, accommodating the genders' differing anatomies, although bikes typically are sold with saddles most appropriate for men. A recumbent bicycle has a reclined chair-like seat that some riders find more comfortable than a saddle, especially riders who suffer from certain types of seat, back, neck, shoulder, or wrist pain. Recumbent bicycles may have either under-seat or over-seat steering. Brakes. Modern bicycle "brakes" may be "rim brakes", in which friction pads are compressed against the wheel rims, "internal hub brakes", in which the friction pads are contained within the wheel hubs, "disc brakes", with a separate rotor for braking. Disc brakes are more common on off-road bicycles, tandems and recumbent bicycles than on road-specific bicycles. With hand-operated brakes, force is applied to brake levers mounted on the handlebars and transmitted via Bowden cables or hydraulic lines to the friction pads. A rear hub brake may be either hand-operated or pedal-actuated, as in the back pedal "coaster brakes" which were popular in North America until the 1960s, and are still common in children's bicycles. Track bicycles do not have dedicated brakes. Brakes are not required for riding on a track because all riders ride in the same direction around a track which does not necessitate sharp deceleration. Track riders are still able to slow down because all track bicycles are fixed-gear, meaning that there is no freewheel. Without a freewheel, coasting is impossible, so when the rear wheel is moving, the crank is moving. To slow down, the rider applies resistance to the pedals this acts as a braking system which can be as effective as a friction-based rear wheel brake, but not as effective as a front wheel brake. Suspension. Bicycle suspension refers to the system or systems used to "suspend" the rider and all or part of the bicycle. This serves two purposes: Bicycle suspensions are used primarily on mountain bicycles, but are also common on hybrid bicycles, and can even be found on some road bicycles, as they can help deal with problematic vibration. Suspension is especially important on recumbent bicycles, since while an upright bicycle rider can stand on the pedals to achieve some of the benefits of suspension, a recumbent rider cannot. Wheels. The wheel axle fits into dropouts in the frame and forks. A pair of wheels may be called a wheelset, especially in the context of ready-built "off the shelf", performance-oriented wheels. Tires vary enormously. Skinny, road-racing tires may be completely smooth, or (slick). On the opposite extreme, off-road tires are much wider and thicker, and usually have a deep tread for gripping in muddy conditions. Accessories, repairs, and tools. Some components, which are often optional accessories on sports bicycles, are standard features on utility bicycles to enhance their usefulness and comfort. Mudguards, or fenders, protect the cyclist and moving parts from spray when riding through wet areas and chainguards protect clothes from oil on the chain while preventing clothing from being caught between the chain and crankset teeth. Kick stands keep a bicycle upright when parked. Front-mounted baskets for carrying goods are often used. Luggage carriers and panniers mounted above the rear tire can be used to carry equipment or cargo. Parents sometimes add rear-mounted child seats and or an auxiliary saddle fitted to the crossbar to transport children. "Toe-clips" and "toestraps" and clipless pedals help keep the foot locked in the proper position on the pedals, and enable the cyclist to pull as well as push the pedals—although not without their hazards, eg. may lock foot in when needed to prevent a fall. Technical accessories include cyclocomputers for measuring speed, distance, etc. Other accessories include lights, reflectors, security locks, mirror, water bottles and cages, and bell. Bicycle helmets may help reduce injury in the event of a collision or accident, and a certified helmet is legally required for some riders in some jurisdictions. Helmets are classified as an accessory or an item of clothing by others. Many cyclists carry "tool kits". These may include a tire patch kit (which, in turn, may contain any combination of a hand pump or CO2 Pump, tire levers, spare tubes, self-adhesive patches, or tube-patching material, an adhesive, a piece of sandpaper or a metal grater (to roughing the tube surface to be patched), and sometimes even a block of French chalk.), wrenches, hex keys, screwdrivers, and a chain tool. There are also cycling specific multi-tools that combine many of these implements into a single compact device. More specialized bicycle components may require more complex tools, including proprietary tools specific for a given manufacturer. Some bicycle parts, particularly hub-based gearing systems, are complex, and many cyclists prefer to leave maintenance and repairs to professional bicycle mechanics. In some areas it is possible to purchase road-side assistance from companies such as the Better World Club. Other cyclists maintain their own bicycles, perhaps as part of their enjoyment of the hobby of cycling or simply for economic reasons. The ability to repair and maintain your own bicycle is also celebrated within the DIY movement. Standards. A number of formal and industry standards exist for bicycle components to help make spare parts exchangeable and to maintain a minimum product safety. The International Organization for Standardization, ISO, has a special technical committee for cycles, TC149, that has the following scope: "Standardization in the field of cycles, their components and accessories with particular reference to terminology, testing methods and requirements for performance and safety, and interchangeability." CEN, European Committee for Standardisation, also has a specific Technical Committee, TC333, that defines European standards for cycles. Their mandate states that EN cycle standards shall harmonize with ISO standards. Some CEN cycle standards were developed before ISO published their standards, leading to strong European influences in this area. European cycle standards tend to describe minimum safety requirements, while ISO standards have historically harmonized parts geometry. Parts. For details on specific bicycle parts, see list of bicycle parts and. Social and historical aspects. The bicycle has had a considerable effect on human society, in both the cultural and industrial realms. In daily life. Around the turn of the 20th century, bicycles reduced crowding in inner-city tenements by allowing workers to commute from more spacious dwellings in the suburbs. They also reduced dependence on horses. Bicycles allowed people to travel for leisure into the country, since bicycles were three times as energy efficient as walking and three to four times as fast. Recently, several European cities have implemented successful schemes known as community bicycle programs or bike-sharing. These initiatives complement a city's public transport system and offer an alternative to motorized traffic to help reduce congestion and pollution. Users take a bicycle at a parking station, use it for a limited amount of time, and then return it to the same or different station. Examples include Bicing in Barcelona, Vélo'v in Lyon and Vélib' in Paris. In cities where the bicycle is not an integral part of the planned transportation system, commuters often use bicycles as elements of a mixed-mode commute, where the bike is used to travel to and from train stations or other forms of rapid transit. Folding bicycles are useful in these scenarios, as they are less cumbersome when carried aboard. Los Angeles removed a small amount of seating on some trains to make more room for bicycles and wheel chairs. Bicycles offer an important mode of transport in many developing countries. Until recently, bicycles have been a staple of everyday life throughout Asian countries. They are the most frequently used method of transport for commuting to work, school, shopping, and life in general. As a result, bicycles there are almost always equipped with baskets. Female emancipation. The diamond-frame safety bicycle gave women unprecedented mobility, contributing to their emancipation in Western nations. As bicycles became safer and cheaper, more women had access to the personal freedom they embodied, and so the bicycle came to symbolize the New Woman of the late 19th century, especially in Britain and the United States. The bicycle was recognized by 19th-century feminists and suffragists as a "freedom machine" for women. American Susan B. Anthony said in a "New York World" interview on February 2 1896: "Let me tell you what I think of bicycling. I think it has done more to emancipate women than anything else in the world. It gives women a feeling of freedom and self-reliance. I stand and rejoice every time I see a woman ride by on a wheel...the picture of free, untrammeled womanhood." In 1895 Frances Willard, the tightly-laced president of the Women’s Christian Temperance Union, wrote a book called "How I Learned to Ride the Bicycle", in which she praised the bicycle she learned to ride late in life, and which she named "Gladys", for its "gladdening effect" on her health and political optimism. Willard used a cycling metaphor to urge other suffragists to action, proclaiming, "I would not waste my life in friction when it could be turned into momentum." Male anger at the freedom symbolized by the New (bicycling) Woman was demonstrated when the male undergraduates of Cambridge University showed their opposition to the admission of women as full members of the university by hanging a woman bicyclist in effigy in the main town square. This was as late as 1897. The bicycle craze in the 1890s also led to a movement for so-called rational dress, which helped liberate women from corsets and ankle-length skirts and other restrictive garments, substituting the then-shocking bloomers. Economic implications. Bicycle manufacturing proved to be a training ground for other industries and led to the development of advanced metalworking techniques, both for the frames themselves and for special components such as ball bearings, washers, and sprockets. These techniques later enabled skilled metalworkers and mechanics to develop the components used in early automobiles and aircraft. They also served to teach the industrial models later adopted, including mechanization and mass production (later copied and adopted by Ford and General Motors), vertical integration (also later copied and adopted by Ford), aggressive advertising (as much as 10% of all advertising in U.S. periodicals in 1898 was by bicycle makers), lobbying for better roads (which had the side benefit of acting as advertising, and of improving sales by providing more places to ride), all first practised by Pope. In addition, bicycle makers adopted the annual model change (later derided as planned obsolescence, and usually credited to General Motors), which proved very successful. Furthermore, bicycles were an early example of conspicuous consumption, being adopted by the fashionable elites. In addition, by serving as a platform for accessories, which could ultimately cost more than the bicycle itself, it paved the way for the likes of the Barbie doll. Moreover, they helped create, or enhance, new kinds of businesses, such as bicycle messengers, travelling seamstresses, riding academies, and racing rinks (Their board tracks were later adapted to early motorcycle and automobile racing.) Also, there were a variety of new inventions, such as spoke tighteners, and specialized lights, socks and shoes, and even cameras (such as the Eastman Company's "Poco"). Probably the best known and most widely used of these inventions, adopted well beyond cycling, is Charles Bennett's Bike Web, which came to be called the "jock strap". They also presaged a move away from public transit that would explode with the introduction of the automobile. This liberation would be repeated again with the appearance of the snowmobile. J. K. Starley's company became the Rover Cycle Company Ltd. in the late 1890s, and then simply the Rover Company when it started making cars. The Morris Motor Company (in Oxford) and Škoda also began in the bicycle business, as did the Wright brothers. Alistair Craig, whose company eventually emerged to become the engine manufacturers Ailsa Craig, also started from manufacturing bicycles, in Glasgow in March 1885. In general, U.S. and European cycle manufacturers used to assemble cycles from their own frames and components made by other companies, although very large companies (such as Raleigh) used to make almost every part of a bicycle (including bottom brackets, axles, etc.) In recent years, those bicycle makers have greatly changed their methods of production. Now, almost none of them produce their own frames. Many newer or smaller companies only design and market their products; the actual production is done by Asian companies. For example, some 60% of the world's bicycles are now being made in China. Despite this shift in production, as nations such as China and India become more wealthy, their own use of bicycles has declined due to the increasing affordability of cars and motorcycles. One of the major reasons for the proliferation of Chinese-made bicycles in foreign markets is the lower cost of labor in China. One of the profound economic implications of bicycle use is that it liberates the user from oil consumption (Ballantine, 1972). H.G. Wells said: “Every time I see an adult on a bicycle, I no longer despair for the future of the human race.” (Quotegarden.com). The bicycle is a inexpensive, fast, healthy and environmentally friendly mode of transport (Illich, 1974) Legal requirements. Early in its development, like in the case of automobiles, there were restrictions on the operation of bicycles. Along with advertising, and to gain free publicity, Albert A. Pope litigated on behalf of cyclists The 1968 Vienna Convention on Road Traffic of the United Nations considers a bicycle to be a vehicle, and a person controlling a bicycle (whether actually riding or not) is considered an operator. The traffic codes of many countries reflect these definitions and demand that a bicycle satisfy certain legal requirements, sometimes even including licensing, before it can be used on public roads. In many jurisdictions, it is an offense to use a bicycle that is not in roadworthy condition. In most jurisdictions, bicycles must have functioning front and rear lights when ridden after dark. As some generator or dynamo-driven lamps only operate while moving, rear reflectors are frequently also mandatory. Since a moving bicycle makes little noise, some countries insist that bicycles have a warning bell for use when approaching pedestrians, equestrians, and other cyclists. See also. "'Special uses and related vehicle types'" References. Other authors: Eddie Borysewicz, Greg LeMond, Davis Phinney, Connie Carpenter.