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top 10 words in brain distribution (in article): cell light animal human form produce body muscle water type |
top 10 words in brain distribution (in article): time type state form century world require term refer provide |
top 10 words in brain distribution (not in article): drink lamp wine beer tissue bottle bone process structure valve |
top 10 words in brain distribution (not in article): plant produce light fruit water drink music grow instrument species |
times more probable under eye 30 20 10 6 4 2.5 1.25 1 1.25 2.5 4 6 10 20 30 times more probable under arch (words not in the model) | |
Eyes'" are organs that detect light, and send signals along the optic nerve to the visual and other areas of the brain. Complex optical systems with resolving power have come in ten fundamentally different forms, and 96% of animal species possess a complex optical system. Image-resolving eyes are present in cnidaria, mollusks, chordates, annelids and arthropods. The simplest "eyes", in even unicellular organisms, do nothing but detect whether the surroundings are light or dark, which is sufficient for the entrainment of circadian rhythms. From more complex eyes, retinal photosensitive ganglion cells send signals along the retinohypothalamic tract to the suprachiasmatic nuclei to effect circadian adjustment. Overview. Complex eyes can distinguish shapes and colors. The visual fields of many organisms, especially predators, involve large areas of binocular vision to improve depth perception; in other organisms, eyes are located so as to maximise the field of view, such as in rabbits and horses. The first proto-eyes evolved among animals 540 million years ago, about the time of the so-called Cambrian explosion. The last common ancestor of animals possessed the biochemical toolkit necessary for vision, and more advanced eyes have evolved in 96% of animal species in 6 of the thirty-something main phyla. In most vertebrates and some mollusks, the eye works by allowing light to enter it and project onto a light-sensitive panel of cells, known as the retina, at the rear of the eye. The cone cells (for color) and the rod cells (for low-light contrasts) in the retina detect and convert light into neural signals for vision. The visual signals are then transmitted to the brain via the optic nerve. Such eyes are typically roughly spherical, filled with a transparent gel-like substance called the vitreous humour, with a focusing lens and often an iris; the relaxing or tightening of the muscles around the iris change the size of the pupil, thereby regulating the amount of light that enters the eye, and reducing aberrations when there is enough light. The eyes of cephalopods, fish, amphibians and snakes usually have fixed lens shapes, and focusing vision is achieved by telescoping the lens — similar to how a camera focuses. Compound eyes are found among the arthropods and are composed of many simple facets which, depending on the details of anatomy, may give either a single pixelated image or multiple images, per eye. Each sensor has its own lens and photosensitive cell(s). Some eyes have up to 28,000 such sensors, which are arranged hexagonally, and which can give a full 360-degree field of vision. Compound eyes are very sensitive to motion. Some arthropods, including many Strepsiptera, have compound eyes of only a few facets, each with a retina capable of creating an image, creating vision. With each eye viewing a different thing, a fused image from all the eyes is produced in the brain, providing very different, high-resolution images. Possessing detailed hyperspectral color vision, the Mantis shrimp has been reported to have the world's most complex color vision system. Trilobites, which are now extinct, had unique compound eyes. They used clear calcite crystals to form the lenses of their eyes. In this, they differ from most other arthropods, which have soft eyes. The number of lenses in such an eye varied, however: some trilobites had only one, and some had thousands of lenses in one eye. In contrast to compound eyes, simple eyes are those that have a single lens. For example, jumping spiders have a large pair of simple eyes with a narrow field of view, supported by an array of other, smaller eyes for peripheral vision. Some insect larvae, like caterpillars, have a different type of simple eye (stemmata) which gives a rough image. Some of the simplest eyes, called ocelli, can be found in animals like some of the snails, which cannot actually "see" in the normal sense. They do have photosensitive cells, but no lens and no other means of projecting an image onto these cells. They can distinguish between light and dark, but no more. This enables snails to keep out of direct sunlight. In organisms dwelling near deep-sea vents, compound eyes have been secondarily simplified and adapted to spot the infra-red light produced by the hot vents in this way the bearers can spot hot springs and avoid being boiled alive. Evolution. Visual pigments appear to have a common ancestor and were probably involved in circadian rhythms or reproductive timing in simple organisms. Complex vision, associated with dedicated visual organs, or eyes, evolved many times in different lineages. Types of eye. Nature has produced ten different eye layouts — indeed every way of capturing an image has evolved at least once in nature, with the exception of zoom and Fresnel lenses. Eye types can be categorized into "simple eyes", with one concave chamber, and "compound eyes", which comprise a number of individual lenses laid out on a convex surface. Note that "simple" does not imply a reduced level of complexity or acuity. Indeed, any eye type can be adapted for almost any behaviour or environment. The only limitations specific to eye types are that of resolution — the physics of compound eyes prevents them from achieving a resolution better than 1°. Also, superposition eyes can achieve greater sensitivity than apposition eyes, so are better suited to dark-dwelling creatures. Eyes also fall into two groups on the basis of their photoreceptor's cellular construction, with the photoreceptor cells either being cilliated (as in the vertebrates) or rhabdomic. These two groups are not monophyletic; the cnidaira also possess cilliated cells, Pit eyes. Pit eyes, also known as stemma, are eye-spots which may be set into a pit to reduce the angles of light that enters and affects the eyespot, to allow the organism to deduce the angle of incoming light. Found in about 85% of phyla, these basic forms were probably the precursors to more advanced types of "simple eye". They are small, comprising up to about 100 cells covering about 100 µm. The directionality can be improved by reducing the size of the aperture, by incorporating a reflective layer behind the receptor cells, or by filling the pit with a refractile material. Pinhole eye. The pinhole eye is an "advanced" form of pit eye incorporating these improvements, most notably a small aperture (which may be adjustable) and deep pit. It is only found in the nautiloids. Without a lens to focus the image, it produces a blurry image, and will blur out a point to the size of the aperture. Consequently, nautiloids can't discriminate between objects with an angular separation of less than 11°. Shrinking the aperture would produce a sharper image, but let in less light. Spherical lensed | An arch'" is a structure that spans a space while supporting weight (e.g. a doorway in a stone wall). Arches appeared as early as the 2nd millennium BC in Mesopotamian brick architecture, but their systematic use started with the Ancient Romans who were the first to apply the technique to a wide range of structures. History. Arches were known by the Mesopotamian, Urartian, Harappan, Egyptian, Babylonian, Greek and Assyrian civilizations, but their use was infrequent and mostly confined to underground structures such as drains where the problem of lateral thrust is greatly diminished. The ancient Romans learned the arch from the Etruscans, refined it and were the first builders to tap its full potential for above ground buildings: "The Romans were the first builders in Europe, perhaps the first in the world, fully to appreciate the advantages of the arch, the vault and the dome." Throughout the Roman empire, their engineers erected arch structures such as bridges, aqueducts, and gates. They also introduced the triumphal arch as a military monument. Vaults began to be used for roofing large interior spaces such as halls and temples, a function which was also assumed by domed structures from the 1st century BC onwards. The Roman arch is semicircular, and built from an odd number of arch bricks (called "voussoirs"). An odd number of bricks is required for there to be a "capstone" or "keystone", the topmost stone in the arch. The Roman arch's shape is the simplest to build, but not the strongest. There is a tendency for the sides to bulge outwards, which must be counteracted by an added weight of masonry to push them inwards. The Romans used this type of semicircular arch freely in many of their secular structures such as aqueducts, palaces and amphitheaters. The semicircular arch was followed in Europe by the pointed Gothic arch or ogive (derived from the Islamic pointed arch in Moorish Spain), whose centreline more closely followed the forces of compression and which was therefore stronger. The semicircular arch can be flattened to make an elliptical arch as in the Ponte Santa Trinita. The parabolic and catenary arches are now known to be the theoretically strongest forms. Parabolic arches were introduced in construction by the Spanish architect Antoni Gaudí, who admired the structural system of Gothic style, but for the buttresses, which he termed “architectural crutches”. The catenary and parabolic arches carry all horizontal thrust to the foundation and so do not need additional elements. The horseshoe arch is based on the semicircular arch, but its lower ends are extended further round the circle until they start to converge. The first examples known are carved into rock in India in the first century AD, while the first known built horseshoe arches are known from Aksum (modern day Ethiopia and Eritrea) from around the 3rd–4th century, around the same time as the earliest contemporary examples in Syria, suggesting either an Aksumite or Syrian origin for the type of arch. It was used in Spanish Visigothic architecture, Islamic architecture and mudéjar architecture, as in the Great Mosque of Damascus and in later Moorish buildings. It was used for decoration rather than for strength. Across the ocean in Mexico and Central America, Mesoamerican civilizations created various types of corbelled arches, such as with the interior tunnels in the Great Pyramid of Cholula and the many styles of corbelled arches built by the Mayan civilization. In Peru, the Inca civilization used a trapezoidal arch in their architecture. The arch is still used today in some modern structures such as bridges. Construction. An arch requires all of its elements to hold it together, raising the question of how an arch is constructed. One answer is to build a frame (historically, of wood) which exactly follows the form of the underside of the arch. This is known as a centre or centring. The voussoirs are laid on it until the arch is complete and self-supporting. For an arch higher than head height, scaffolding would in any case be required by the builders, so the scaffolding can be combined with the arch support. Occasionally arches would fall down when the frame was removed if construction or planning had been incorrect. (The A85 bridge at Dalmally, Scotland suffered this fate on its first attempt, in the 1940s). The interior and lower line or curve of an arch is known as the "intrados". Old arches sometimes need reinforcement due to decay of the keystones, known as bald arch. The gallery shows arch forms displayed in roughly the order in which they were developed. Technical aspects. The arch is significant because, in theory at least, it provides a structure which eliminates tensile stresses in spanning an open space. All the forces are resolved into compressive stresses. This is useful because several of the available building materials such as stone, cast iron and concrete can strongly resist compression but are very weak when tension, shear or torsional stress is applied to them. By using the arch configuration, significant spans can be achieved. This is because all the compressive forces hold it together in a state of equilibrium. This even applies to frictionless surfaces. However, one downside is that an arch pushes outward at the base, and this needs to be restrained in some way, either with heavy sides and friction or angled cuts into bedrock or similar. This same principle holds when the force acting on the arch is not vertical such as in spanning a doorway, but horizontal, such as in arched retaining walls or dams. Even when using concrete, where the structure may be monolithic, the principle of the arch is used so as to benefit from the concrete's strength in resisting compressive stress. Where any other form of stress is raised, it has to be resisted by carefully placed reinforcement rods or fibres. (See Arch bridge.) Other types. The Delicate Arch, a natural arch in Moab, UtahA blind arch is an arch infilled with solid construction so it cannot function as a window, door, or passageway. A dome is a three-dimensional application of the arch, rotated about the center axis. Igloos are notable vernacular structures making use of domes. Natural rock formations may also be referred to as arches. These natural arches are formed by erosion rather than being carved or constructed by man. See Arches National Park for examples. A special form of the arch is the triumphal arch, usually built to celebrate a victory in war. A famous example is the Arc de Triomphe in Paris, France. A vault is an application of the arch extended horizontally in two dimensions; the groin vault is the intersection of two vaults. |