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tomato |
top 10 words in brain distribution (in article): species animal male breed female human hunt wolf live food |
top 10 words in brain distribution (in article): species century form time style type term ride culture modern |
top 10 words in brain distribution (not in article): bird cat egg wild dog elephant pet deer water ant |
top 10 words in brain distribution (not in article): wear horse woman clothe saddle tea fashion dress body design |
times more probable under bear 30 20 10 6 4 2.5 1.25 1 1.25 2.5 4 6 10 20 30 times more probable under tomato (words not in the model) | |
Bears'" are mammals of the family "'Ursidae'". Bears are classified as caniforms, or doglike carnivorans, with the pinnipeds being their closest living relatives. Although there are only eight living species of bear, they are widespread, appearing in a wide variety of habitats throughout the Northern Hemisphere and partially in the Southern Hemisphere. That which pertains to bears is called "ursine". Bears are found in the continents of North America, South America, Europe, and Asia. Common characteristics of modern bears include a large body with stocky legs, a long snout, shaggy hair, plantigrade paws with five nonretractile claws, and a short tail. While the polar bear is mostly carnivorous and the giant panda feeds almost entirely on bamboo, the remaining six species are omnivorous, with largely varied diets including both plants and animals. With the exceptions of courting individuals and mothers with their young, bears are typically solitary animals. They are sometimes diurnal, but are usually active during the night (nocturnal) or twilight (crepuscular). Bears are aided by an excellent sense of smell, and despite their heavy build and awkward gait, they can run quickly and are adept climbers and swimmers. In autumn some bear species forage large amounts of fermented fruits which affects their behaviour.Bears use shelters such as caves and burrows as their dens, which are occupied by most species during the winter for a long period of sleep similar to hibernation. Bears have been hunted since prehistoric times for their meat and fur. To this day, they play a prominent role in the arts, mythology, and other cultural aspects of various human societies. In modern times, the bear's existence has been pressured through the encroachment of their habitats and the illegal trade of bears and bear parts, including the Asian bile bear market. The IUCN lists six bear species as vulnerable or endangered, and even "least concern" species such as the brown bear are at risk of extirpation in certain countries. The poaching and international trade of these most threatened populations is prohibited, but still ongoing. Evolutionary relationships. Fossil of Cave bear ("Ursus spelaeus") The Ursidae family belongs to the order Carnivora and is one of nine families in the suborder Caniformia, or "doglike" carnivorans. Bears' closest living relatives are the pinnipeds, a clade of three families: Odobenidae (the walrus), Otariidae (fur seals and sea lions), and Phocidae (true or earless seals). Bears comprise eight species in three subfamilies: Ailuropodinae (monotypic with the giant panda), Tremarctinae (monotypic with the Spectacled Bear), and Ursinae (containing six species divided into one to three genera, depending upon authority). The origins of Ursidae can be traced back to the very small and graceful "Parictis" that had a skull only 7 cm (3 in) long. Parictis first occur in North America in the Late Eocene (ca. 38 million years ago), but this genus did not appear in Eurasia and Africa until the Miocene. The raccoon-sized, dog-like "Cephalogale", however, is widely regarded as the most primitive ursid and is ideally suited as a representative basal taxon for the family. "Cephalogale" first appeared during the middle Oligocene and early Miocene (approximately 20–30 million years ago) in Europe. "Cephalogale" gave rise to a lineage of early bears of the genus "Ursavus". This genus radiated in Asia and ultimately gave rise to the first true bears (genus "Ursus") in Europe, 5 million years ago. Even among its primitive species, such as "C. minor", it exhibits typical ursid synapomorphic dentition such as posteriorly oriented M2 postprotocrista molars, elongated m2 molars, and a reduction of the premolars. Living members of the ursids are morphologically well defined by their hypocarnivorous (non-strictly meat-eating) dentitions, but fossil ursids include hypercarnivorous (strictly meat-eating) taxa, although they never achieved the extreme hypercarnivory seen in mustelids. Cephalogale was a mesocarnivore (intermediate meat-eater). Other extinct bear genera include "Arctodus", "Agriarctos", "Plionarctos" and "Indarctos". It is uncertain whether ursids were in Asia during the late Eocene, although there is some suggestion that a limited immigration from Asia may have produced "Parictis" in North America due to the major sea level lowstand at ca. 37 Ma, but no "Parictis" fossils have yet to be found in East Asia. Ursids did, however, become very diversified in Asia later during the Oligocene. Four genera representing two subfamilies (Amphicynodontinae and Hemicyoninae) have been discovered in the Oligocene of Asia: "Amphicticeps", "Amphicynodon", "Pachycynodon", and "Cephalogale". "Amphicticeps" is endemic from Asia and the other three genera are common to both Asia and Europe. This indicates migration of ursids between Asia and Europe during the Oligocene and migration of several taxa from Asia to North America likely occurred later during the late Oligocene or early Miocene. Although "Amphicticeps" is morphologically closely related to "Allocyon", and also to "Kolponomos" of North America, no single genus of the Ursidae from this time period is known to be common to both Eurasia and North America. Cephalogale, however, do appear in North America in the early Miocene. It is interesting to note that rodents, such as "Haplomys" and "Pseudotheridomys" (late Oligocene) and "Plesiosminthus" and "Palaeocastor" (early Miocene), are common to both Asia and North America and this indicates that faunal exchange did occur between Asia and North America during the late Oligocene to early Miocene. Ursid migration from Asia to North America would therefore have also been very likely to occur during this time. In the late Neogene three major carnivoran migrations that definitely included ursids are recognized between Eurasia and North America. The first (probably 21–18 Ma) was waves of intermittent dispersals including "Amphicynodon", "Cephalogale" and "Ursavus". The second migration occurred about 7–8 Ma and included "Agriotherium" – this was unusual among ursoids in that it also colonised sub-Saharan Africa. The third wave took place in the early Pliocene 4 Ma, consisting of "Ursus". The giant panda's taxonomy has long been debated. Its original classification by Armand David in 1869 was within the bear genus Ursus, but in 1870 it was reclassified by Alphonse Milne-Edwards to the raccoon family. In recent studies, the majority of DNA analyses suggest that the giant panda has a much closer relationship to other bears and should be considered a member of the family Ursidae. The status of the red panda remains uncertain, but many experts, including Wilson and Reeder, classify it as a member of the bear family. Others place it with the raccoons in Procyonidae or in its own family, the Ailuridae. Multiple similarities between the two pandas, including the presence of false thumbs, are thought to represent convergent evolution for feeding primarily on bamboo. There is also evidence that, unlike their neighbors elsewhere, the brown bears of Alaska's ABC islands are more closely related to polar bears than they are to other brown bears in the world. Researchers Gerald Shields and Sandra Talbot of the University of Alaska Fairbanks Institute of Arctic Biology studied the DNA of several samples of the species and found that their DNA is different from that of other brown bears. The researchers discovered that their DNA was unique compared to brown bears anywhere else in the world. The discovery has shown that while all other brown bears share a brown bear as their closest relative, those of Alaska's ABC Islands differ and share their closest relation with the polar bear. There is also supposed to be a very rare large bear in China called the blue bear, which presumably is a type of black bear. This animal has never been photographed. Koalas are often referred to as bears due to their appearance; they are not bears, however, but marsupials. Classification. The genera "Melursus" and "Helarctos" are sometimes also included in "Ursus". The Asiatic black bear and the polar bear used to be placed in their own genera, "Selenarctos" and "Thalarctos" which are now placed at subgenus rank. A number of hybrids have been bred between American black, brown, and polar bears (see Ursid hybrids). Biology. Despite being quadrupeds, bears can stand and sit similarly to humans. Bears are generally bulky and robust animals with relatively short legs. Unlike other land carnivorans, bears stand and walk on the soles of their feet rather than on their toes. They distribute their weight toward the hindfeet which makes then look lumbering when they walk. They are still quite fast with the brown bear reaching 30 mph although they are still slower than felines and canines. Bear can stand on their hindfeet and sit up straight with remarkable balance. Bears have non-retractable claws which are used for digging, climbing, tearing and catching prey. Their ears are rounded. Dentition. Unlike most other members of the Carnivora, bears have relatively undeveloped carnassial teeth, and their teeth are adapted for a diet that includes a significant amount of vegetable matter. The canine teeth are large, and the molar teeth flat and crushing. There is considerable variation in dental formula even within a given species. It has been suggested that this indicates bears are still in the process of evolving from a carnivorous to a predominantly herbivorous diet. Polar bears appear to have secondarily re-evolved fully functional carnassials, as their diet has switched back towards carnivory. The dental formula for living bears is: Diet & Interspecific Interactions. Their carnivorous reputation non-withstanding, most bears have adopted to a diet comprised of more plant than animal matter and are completely opportunistic omnivores. One exception is the Polar Bear, who has had to adopt a diet of mainly marine mammals to survive in the Arctic. The other exception is the Giant Panda has adapted a diet comprised mainly of bamboo. The Sloth Bear, though not as specialized as the previous two species, has lost several front teeth usually seen in bears and developed a long, suctioning tongue in order to feed on the termites and other burrowing insects that they favor. All bears will feed on any food source that becomes available. When taking warm-blooded animals, bears will typically take small or young animals, because of the endurance and potential danger that comes with attacking large prey. Although (besides Polar Bears) both species of black bear and the Brown Bear can sometimes take large prey, such as ungulates. Often, bears will feed on other large animals when they encounter a carcass, whether or not the carcass is claimed by or is the kill of another predator. This competition is the main source of interspecies conflict. Bears are typically the apex predators in their range due to their size and power, and can defend a carcass against nearly all comers. Mother bears also can usually defend their cubs against other predators. The Tiger is the only known predator known to regularly prey on adult bears, including Sloth Bears, Asiatic Black Bears, Giant Pandas, Sun Bears and small Brown Bears. Reproduction. The bear's courtship period is very brief. Bears in northern climates reproduce seasonally, usually after a period of inactivity similar to hibernation, although tropical species breed all year round. Cubs are born toothless, blind, and bald. The cubs of brown bears, usually born in litters of 1–3, will typically stay with the mother for two full seasons. They feed on their mother's milk through the duration of their relationship with their mother, although as the cubs continue to grow, nursing becomes less frequent and learn to begin hunting with the mother. They will remain with the mother for approximately three years, until she enters the next cycle of estrus and drives the cubs off. Bears will reach sexual maturity in five to seven years. Male bears, especially Polar and Brown Bears, will kill and sometimes devour cubs born to another father in order to induce a female to breed again. Female bears are often successful in driving off males in protection of their cubs, despite being rather smaller. Winter dormancy. Many bears of northern regions are assumed to hibernate in the winter. While many bear species do go into a physiological state called hibernation or winter sleep, it is not true hibernation. In true hibernators, body temperatures drop to near ambient and heart rate slows drastically, but the animals periodically rouse themselves to urinate or defecate and to eat from stored food. The body temperature of bears, on the other hand, drops only a few degrees from normal and heart rate slows only slightly. They normally do not wake during this "hibernation", and therefore do not eat, drink, urinate or defecate the entire period. Higher body heat and being easily roused may be adaptations, because females give birth to their cubs during this winter sleep. It can therefore be considered a more efficient form of hibernation because they need not awake through the entire period, but they are more quickly and easily awakened at the end of their hibernation. They have to stay in a den for the whole hibernation. Relationship with humans. Some species, such as the polar bear, American black bear, Sloth Bear and the brown bear, are dangerous to humans, especially in areas where they have become used to people. On the west coast of Canada, the American black bear has become an integral part of the silviculture industries, specifically treeplanting. The bears are coaxed into areas of harvested forest to "flush out" the other wildlife, i.e. moose, which are a far greater threat to planters. All bears are physically powerful and are likely capable of fatally attacking a person, but they, for the most part, are shy, easily frightened and will avoid humans. The danger that bears pose is often vastly exaggerated, in part by the human imagination. However, when a mother feels her cubs are threatened, she will behave ferociously. It is recommended to give all bears a wide berth because they are behaviorally unpredictable. Laws have been passed in many areas of the world to protect bears from hunters habitat destruction. Some populated areas with bear populations have also outlawed the feeding of bears, including allowing them access to garbage or other food waste. Bears in captivity have been trained to dance, box, or ride bicycles; however, this use of the animals became controversial in the late 20th century. Bears were kept for baiting in Europe at least since the 16th century. Bears as food and medicine. Many people enjoy hunting bears and eating them. Their meat is dark and stringy, like a tough cut of beef. In Cantonese cuisine, bear paws are considered a delicacy. The peoples of China, Japan, and Korea use bears' body parts and secretions (notably their gallbladders and bile) as part of traditional Chinese medicine. It is believed more than 12,000 bile bears are kept on farms, farmed for their bile, in China, Vietnam and South Korea. Bear meat must be cooked thoroughly as it can often be infected with trichinellosis. Myth and legend. Some evidence has been brought to light on prehistoric bear worship, see Arctic, Arcturus, Great Bear, Berserker, Kalevala. Anthropologists such as Joseph Campbell have regarded this as a common feature in most of the fishing and hunting-tribes. The prehistoric Finns, along with most Finno-Ugric peoples, considered the bear as the spirit of one's forefathers. This is why the bear was a greatly respected animal, with several euphemistic names. The bear is the national animal of Finland. This kind of attitude is reflected in the traditional Russian fairy tale "Morozko", whose arrogant protagonist Ivan tries to kill a mother bear and her cubs — and is punished and humbled by having his own head turned magically into a bear's head and being subsequently shunned by human society. "The Brown Bear of Norway" is a Scottish fairy tale telling the adventures of a girl who married a prince magically turned into a bear, and who managed to get him back into a human form by the force of her love and after many trials and difficulties. There has been evidence about early bear worship in China and among the Ainu culture as well (see Iomante). Korean people in their mythology identify the bear as their ancestor and symbolic animal. According to the Korean legend, a god imposed a difficult test on a she-bear, and when she passed it the god turned her into a woman and married her. In addition, the Proto-Indo-European word for bear, "*h₂ŕ̥tḱos" (ancestral to the Greek "arktos", Latin "ursus", Welsh "arth" (cf. Arthur), Albanian ari, Armenian arj, Sanskrit "ṛkṣa", Hittite "ḫartagga") seems to have been subject to taboo deformation or replacement (as was the word for wolf, "wlkwos"), resulting in the use of numerous unrelated words with meanings like "brown one" (English "bruin") and "honey-eater" (Slavic "medved"). Thus four Indo-European language groups do not share the same PIE root. The theory of the bear taboo is taught to almost all beginning students of Indo-European and historical linguistics; the putative original PIE word for bear is itself descriptive, because a cognate word in Sanskrit is "rakṣas", meaning "harm, injury". Legends of saints taming bears are common in the Alpine zone. In the arms of the bishopric of Freising ("see illustration") the bear is the dangerous totem animal tamed by St. Corbinian and made to carry his civilised baggage over the mountains. A bear also features prominently in the legend of St. Romedius, who is also said to have tamed one of these animals and had the same bear carry him from his hermitage in the mountains to the city of Trento. Similar stories are told of Saint Gall and Saint Columbanus. This recurrent motif was used by the Church as a symbol of the victory of Christianity over Paganism, represented by the fiery. Imaginary bears are a popular feature of many children's stories including Goldilocks and the Three Bears, the Berenstein Bears, and Winnie the Pooh. The constellations Ursa Major and Ursa Minor represent bears. Symbolic use. The Russian bear is a common National personification for Russia (as well as the Soviet Union) and even Germany. The brown bear is Finland's national animal. In the United States, the black bear is the state animal of Louisiana, New Mexico, and West Virginia; the grizzly bear is the state animal of both Montana and California. Bears appear in the canting arms of Berne and Berlin. Also, "bear", "bruin", or specific types of bears are popular nicknames or mascots, e.g. for sports teams (Chicago Bears, Boston Bruins); and a bear cub called Misha was mascot of the 1980 Summer Olympics in Moscow, USSR. Smokey Bear has become a part of American culture since his introduction in 1944. Known to almost all Americans, he and his message, "Only You Can Prevent Forest Fires" (updated in 2001 to "Only You Can Prevent Wildfires") has been a symbol of preserving woodlands. Smokey wears a hat similar to one worn by many U.S. state police officers, giving rise to the CB slang "bear" or "Smokey" for the highway patrol. Figures of speech. The physical attributes and behaviours of bears are commonly used in figures of speech in English. Teddy bears. Around the world, many children have stuffed animals in the form of bears. Names. In Scandinavia the word for bear is "Björn" (or "Bjørn"), and is a relatively common given name for males. The use of this name is ancient and has been found mentioned in several runestone inscriptions. The name was also used by J.R.R. Tolkien in his book "The Hobbit", where a bear-like character is named Beorn. The female first name "Ursula", originally derived from a Christian saint's name and common in English- and German-speaking countries, means "Little she-bear" (dimunitive of Latin "ursa"). In Switzerland the male first name "Urs" is especially popular. In Russian and other Slavic languages, the word for bear, "Medved" (медведь), and variants or derivatives such as Medvedev are common surnames. The Irish family name "McMahon" means "Son of Bear" in Irish. One of widely held etymological explanations for the common name "Arthur" is that it originally meant "bear-like". In East European Jewish communities, the name "Ber" (בער) — Yiddish cognate of "Bear" — has been attested as a common male first name, at least since the 18th century, and was among others the name of several prominent Rabbis. The Yiddish "Ber" is still in use among Orthodox Jewish communities in Israel, the US and other countries. With the transition from Yiddish to Hebrew under the influence of Zionism, the Hebrew word for "bear", "Dov" (דב), was taken up in contemporary Israel and is at present among the commonly used male first names in that country. "Ten Bears" (Paruasemana) was the name of a well-known 19th Century chieftain among the Comanche. Also among other Native American tribes, bear-related names are attested. | The Tomato'" ("Solanum lycopersicum", syn. "Lycopersicon lycopersicum" & "Lycopersicon esculentum") is an herbaceous, usually sprawling plant in the Solanaceae or nightshade family, as are its close cousins tobacco, potatoes, aubergine (eggplants), chilli peppers, and the poisonous belladonna. It is a perennial, often grown outdoors in temperate climates as an annual. Typically reaching to 1-3m (3 to 9 ft) in height, it has a weak, woody stem that often vines over other plants. The leaves are long, odd pinnate, with 5–9 leaflets on petioles, each leaflet up to long, with a serrated margin; both the stem and leaves are densely glandular-hairy. The flowers are across, yellow, with five pointed lobes on the corolla; they are borne in a cyme of 3–12 together. The tomato is native to South America and a prehistoric introduction to Central America and Southern parts of North America. Genetic evidence shows that the progenitors of tomatoes were herbaceous green plants with small green fruit with a center of diversity in the highlands of Peru. These early Solanums diversified into the ~dozen species of tomato recognized today. One species, Solanum Lycopersicum, was transported to Mexico where it was grown and consumed by prehistoric humans. The exact date of domestication is not known. Evidence supports the theory the first domesticated tomato was a little yellow fruit, ancestor of L. cerasiforme, grown by the Aztecs of Central Mexico who called it ‘xitomatl’ (pronounced zee-toe-má-tel), meaning plump thing with a navel, and later called tomatl by other Mesoamerican peoples. Aztec writings mention tomatoes were prepared with peppers, corn and salt, likely to be the original salsa recipe. Some believe Spanish explorer Cortez may have been the first to transfer the small yellow tomato to Europe after he captured the Aztec city of Tenochtítlan in 1521, now Mexico City. Yet others believe Christopher Columbus, an Italian working for the Spanish monarchy, discovered the tomato earlier in 1493. The earliest discussion of the tomato in European literature appeared in an herbal written in 1544 by Pietro Andrea Mattioli, an Italian physician and botanist, who named it pomi d’oro, golden apple. The word "tomato" comes from a word in the Nahuatl language, "tomatl". The specific name, "lycopersicum", means "wolf-peach" (compare the related species "Solanum lycocarpum", whose scientific name means "wolf-fruit", common name "wolf-apple"), as they are a major food of wild canids in South America. Early history. Two modern tomato cultivar groups, one represented by the Matt's Wild Cherry tomato, the other by currant tomatoes, originate by recent domestication of the wild tomato plants apparently native to eastern Mexico. Aztecs and other peoples in the region used the fruit in their cooking; it was being cultivated in southern Mexico and probably other areas by 500BC. It is thought that the Pueblo people believed that those who witnessed the ingestion of tomato seeds were blessed with powers of divination. The large, lumpy tomato, a mutation from a smoother, smaller fruit, originated and was encouraged in Mesoamerica. Smith states this variant is the direct ancestor of some modern cultivated tomatoes. According to Andrew F Smith's "The Tomato in America", the tomato probably originated in the highlands of the west coast of South America. Smith notes there is no evidence the tomato was cultivated or even eaten in Peru before the Spanish arrived. Spanish distribution. After the Spanish colonization of the Americas, the Spanish distributed the tomato throughout their colonies in the Caribbean. They also took it to the Philippines, whence it moved to southeast Asia and then the entire Asian continent. The Spanish also brought the tomato to Europe. It grew easily in Mediterranean climates, and cultivation began in the 1540s. It was probably eaten shortly after it was introduced, and was certainly being used as food by the early 1600s in Spain. The earliest discovered cookbook with tomato recipes was published in Naples in 1692, though the author had apparently obtained these recipes from Spanish sources. However, in certain areas of Italy, such as Florence, the fruit was used solely as tabletop decoration before it was incorporated into the local cuisine in the late 17th or early 18th century. In Britain. Tomatoes were not grown in England until the 1590s, according to Smith. One of the earliest cultivators was John Gerard, a barber-surgeon. Gerard's "Herbal", published in 1597 and largely plagiarized from continental sources, is also one of the earliest discussions of the tomato in England. Gerard knew that the tomato was eaten in Spain and Italy. Nonetheless, he believed that it was poisonous (tomato leaves and stems actually contain poisonous glycoalkaloids, but the fruit is safe). Gerard's views were influential, and the tomato was considered unfit for eating (though not necessarily poisonous) for many years in Britain and its North American colonies. But by the mid-1700s, tomatoes were widely eaten in Britain; and before the end of that century, the "Encyclopædia Britannica" stated that the tomato was "in daily use" in soups, broths, and as a garnish. In Victorian times, cultivation reached an industrial scale in glasshouses, most famously in Worthing. Pressure for housing land in the 1930s to 1960s saw the industry move west to Littlehampton, and to the market gardens south of Chichester. Over the past 15 years, the British tomato industry has declined as more competitive imports from Spain and the Netherlands have reached the supermarkets. North America. The earliest reference to tomatoes being grown in British North America is from 1710, when herbalist William Salmon reported seeing them in what is today South Carolina. They may have been introduced from the Caribbean. By the mid-18th century, they were cultivated on some Carolina plantations, and probably in other parts of the Southeast as well. It is possible that some people continued to think tomatoes were poisonous at this time; and in general, they were grown more as ornamental plants than as food. Thomas Jefferson, who ate tomatoes in Paris, sent some seeds back to America. Because of their longer growing season for this heat-loving crop, several states in the US Sun Belt became major tomato-producers, particularly Florida and California. In California tomatoes are grown under irrigation for both the fresh fruit market and for canning and processing. The University of California, Davis (UC Davis) became a major center for research on the tomato. The C.M. Rick Tomato Genetics Resource Center at UC Davis is a genebank of wild relatives, monogenic mutants and miscellaneous genetic stocks of tomato. The Center is named for the late Dr. Charles M. Rick, a pioneer in tomato genetics research. Research on processing tomatoes is also conducted by the in Escalon, CA. Production trends. 125 million tons of tomatoes were produced in the world in 2005. China, the largest producer, accounted for about one quarter of the global output, followed by United States and Turkey. For processing tomatoes, California accounts for 90% of US production and 35% of world production. According to FAOSTAT, the top producers of tomatoes (in tonnes) in 2005 were: Cultivation and uses. The tomato is now grown worldwide for its edible fruits, with thousands of cultivars having been selected with varying fruit types, and for optimum growth in differing growing conditions. Cultivated tomatoes vary in size from cherry tomatoes, about the same 1–2 cm size as the wild tomato, up to beefsteak tomatoes 10 cm or more in diameter. The most widely grown commercial tomatoes tend to be in the 5–6 cm diameter range. Most cultivars produce red fruit; but a number of cultivars with yellow, orange, pink, purple, green, black, or white fruit are also available. Multicolored and striped fruit can also be quite striking. Tomatoes grown for canning are often elongated, 7–9 cm long and 4–5 cm diameter; they are known as plum tomatoes. Roma-type tomatoes are important cultivars in the Sacramento Valley where a 120-acre Morning Star cannery handles 1.2 million pounds of tomatoes an hour during the harvest season where the fields yield about 40 tons to the acre. Tomatoes are one of the most common garden fruits in the United States and, along with zucchini, have a reputation for outproducing the needs of the grower. As in most sectors of agriculture, there is increasing demand in developed countries for organic tomatoes, as well as heirloom tomatoes, to make up for flavor and texture faults in commercial tomatoes. Quite a few seed merchants and banks provide a large selection of heirloom seeds. Tomato seeds are occasionally organically produced as well, but only a small percentage of organic crop area is grown with organic seed. The definition of a heirloom tomato is vague, but unlike commercial hybrids, all are self-pollinators who have bred true for 40 years or more. Varieties. There are a great many (around 7500) tomato varieties grown for various purposes. Heirloom strains are becoming increasingly popular, particularly among home gardeners and organic producers, since they tend to produce more interesting and flavorful crops at the cost of disease resistance, and productivity. Hybrid plants remain common, since they tend to be heavier producers and sometimes combine unusual characteristics of heirloom tomatoes with the ruggedness of conventional commercial tomatoes. Tomato varieties are roughly divided into several categories, based mostly on shape and size. "Slicing" or "globe" tomatoes are the usual tomatoes of commerce; beefsteak are large tomatoes often used for sandwiches and similar applications- their kidney-bean shape makes commercial use impractical along with a thinner skin and being not bred for a long shelf life; globe tomatoes are of the category of canners used for a wide variety of processing and fresh eating; oxheart tomatoes can range in size up to beefsteaks, and are shaped like large strawberries; plum tomatoes, or paste tomatoes (including pear tomatoes), are bred with a higher solid content for use in tomato sauce and paste and are usually oblong; pear tomatoes are obviously pear shaped and based upon the San Marzano types for a richer gourmet paste; cherry tomatoes are small and round, often sweet tomatoes generally eaten whole in salads; and grape tomatoes which are a more recent introduction are smaller and oblong used in salads. Tomatoes are also commonly classified as determinate or indeterminate. Determinate, or bush, types bear a full crop all at once and top off at a specific height; they are often good choices for container growing. Determinate types are preferred by commercial growers who wish to harvest a whole field at one time, or home growers interested in canning. Indeterminate varieties develop into vines that never top off and continue producing until killed by frost. They are preferred by home growers and local-market farmers who want ripe fruit throughout the season. As an intermediate form, there are plants sometimes known as "vigorous determinate" or "semi-determinate"; these top off like determinates but produce a second crop after the initial crop. The majority of heirloom tomatoes are indeterminate, although some determinate heirlooms exist. Most modern tomato cultivars are smooth surfaced but some older tomato cultivars and most modern beefsteaks often show pronounced ribbing, a feature that may have been common to virtually all pre-Columbian cultivars. While virtually all commercial tomato varieties are red, some tomato cultivars, especially heirlooms, produce fruit in colors other than red, including yellow, orange, pink, black, brown, ivory, white, and purple, though such fruit is not widely available in grocery stores, nor are their seedlings available in typical nurseries, but must be bought as seed, often via mail-order. Less common variations include fruit with stripes (Green Zebra), fuzzy skin on the fruit (Fuzzy Peach, Red Boar), multiple colors (Hillbilly, Burracker's Favorite, Lucky Cross), etc. There is also a considerable gap between commercial and home-gardener cultivars; home cultivars are often bred for flavor to the exclusion of all other qualities, while commercial cultivars are bred for such factors as consistent size and shape, disease and pest resistance, and suitability for mechanized picking and shipping. Tomatoes grow well with 7 hours of sunlight a day. A fertilizer with the ratio 5-10-10 can be used for extra growth, but manure or compost works well too. Diseases and pests. Tomato cultivars vary widely in their resistance to disease. Modern hybrids focus on improving disease resistance over the heirloom plants. One common tomato disease is tobacco mosaic virus, and for this reason smoking or use of tobacco products are discouraged around tomatoes, although there is some scientific debate over whether the virus could possibly survive being burned and converted into smoke. Various forms of mildew and blight are also common tomato afflictions, which is why tomato cultivars are often marked with a combination of letters which refer to specific disease resistance. The most common letters are: V'" -"verticillium" wilt, F'" -"fusarium" wilt strain I, FF'" -"fusarium" wilt strain I & II, N'" -"nematodes", T'" -"tobacco mosaic virus", and A'" -"alternaria". Another particularly dreaded disease is curly top, carried by the beet leafhopper, which interrupts the lifecycle, ruining a nightshade plant as a crop. As the name implies, it has the symptom of making the top leaves of the plant wrinkle up and grow abnormally. Some common tomato pests are cutworms, tomato hornworms and tobacco hornworms, aphids, cabbage loopers, whiteflies, tomato fruitworms, flea beetles, red spider mite, slugs, and Colorado potato beetles. Pollination. In the wild, original state, tomatoes required cross-pollination; they were much more self-incompatible than domestic cultivars. As a floral device to reduce selfing, the pistils of wild tomatoes extended farther out of the flower than today's cultivars. The stamens were, and remain, entirely within the closed corolla. As tomatoes were moved from their native areas, their traditional pollinators, (probably a species of halictid bee) did not move with them. The trait of self-fertility (or self-pollenizing) became an advantage and domestic cultivars of tomato have been selected to maximize this trait. This is not the same as self-pollination, despite the common claim that tomatoes do so. That tomatoes pollinate themselves poorly without outside aid is clearly shown in greenhouse situations where pollination must be aided by artificial wind, vibration of the plants (one brand of vibrator is a wand called an "electric bee" that is used manually), or more often today, by cultured bumblebees. The anther of a tomato flower is shaped like a hollow tube, with the pollen produced within the structure rather than on the surface, as with most species. The pollen moves through pores in the anther, but very little pollen is shed without some kind of outside motion. The best source of outside motion is a sonicating bee such as a bumblebee or the original wild halictid pollinator. In an outside setting, wind or biological agents provide sufficient motion to produce commercially viable crops. Hydroponic and greenhouse cultivation. Tomatoes are often grown in greenhouses in cooler climates, and indeed there are cultivars such as the British 'Moneymaker' and a number of cultivars grown in Siberia that are specifically bred for indoor growing. In more temperate climates, it is not uncommon to start seeds in greenhouses during the late winter for future transplant. With the transplanting of tomatoes, there is a process of hardening that the plant must go through before being able to be placed outside in order to have greater survival. Hydroponic tomatoes are also available, and the technique is often used in hostile growing environments as well as high-density plantings. Picking and ripening. Tomatoes are often picked unripe (and thus colored green) and ripened in storage with ethylene. Unripe tomatoes are firm, as they ripen they soften until reaching the ripe state where they are red or orange in color and slightly soft to the touch. Ethylene is a hydrocarbon gas produced by many fruits that acts as the molecular cue to begin the ripening process. Tomatoes ripened in this way tend to keep longer but have poorer flavor and a mealier, starchier texture than tomatoes ripened on the plant. They may be recognized by their color, which is more pink or orange than the other ripe tomatoes' deep red. In 1994 Calgene introduced a genetically modified tomato called the 'FlavrSavr' which could be vine ripened without compromising shelf life. However, the product was not commercially successful (see main article for details) and was only sold until 1997. Recently, stores have begun selling "tomatoes on the vine", which are determinate varieties that are ripened or harvested with the fruits still connected to a piece of vine. These tend to have more flavor than artificially ripened tomatoes (at a price premium), but still may not be the equal of local garden produce. Slow-ripening cultivars of tomato have been developed by crossing a non-ripening cultivar with ordinary tomato cultivars. Cultivars were selected whose fruits have a long shelf life and at least reasonable flavor. Modern uses and nutrition. Tomatoes are now eaten freely throughout the world, and their consumption is believed to benefit the heart among other things. They contain lycopene, one of the most powerful natural antioxidants, which, especially when tomatoes are cooked, has been found to help prevent prostate cancer. However, other research contradicts this claim. Tomato extract branded as Lycomato is now also being promoted for treatment of high blood pressure. Lycopene has also been shown to improve the skin's ability to protect against harmful UV rays. Natural genetic variation in tomatoes and their wild relatives has given a genetic treasure trove of genes that produce lycopene, carotene, anthocyanin, and other antioxidants. Tomato varieties are available with double the normal vitamin C (Doublerich), 40 times normal vitamin A (97L97), high levels of anthocyanin (P20 Blue), and two to four times the normal amount of lycopene (numerous available cultivars with the high crimson gene). Though it is botanically a berry, a subset of fruit, the tomato is nutritionally categorized as a vegetable (see below). Since "vegetable" is not a botanical term, there is no contradiction in a plant part being a fruit botanically while still being considered a vegetable. Tomatoes are used extensively in Mediterranean cuisine, especially Italian and Middle Eastern cuisines. The tomato is acidic; this acidity makes tomatoes especially easy to preserve in home canning whole, in pieces, as tomato sauce, or paste. Tomato juice is often canned and sold as a beverage; Unripe green tomatoes can also be breaded and fried, used to make salsa, or pickled. Cultural impact===. The town of Buñol, Spain, annually celebrates La Tomatina, a festival centered on an enormous tomato fight. Tomatoes are also a popular "non-lethal" throwing weapon in mass protests; and there was a common tradition of throwing rotten tomatoes at bad performers on a stage during the 19th century; today it is usually referenced as a mere metaphor (see Rotten Tomatoes). Embracing it for this protest connotation, the Dutch Socialist party adopted the tomato as their logo. Known for its tomato growth and production, the Mexican state of Sinaloa takes the tomato as its symbol. Storage. Most tomatoes today are picked before fully ripened. They are bred to continue ripening, but the enzyme that ripens tomatoes stops working when it reaches temperatures below 12.5°C (54.5°F). Once an unripe tomato drops below that temperature, it will not continue to ripen. Once fully ripe, tomatoes can be stored in the refrigerator but are best kept a room temperature. Tomatoes stored in the refrigerator tend to lose flavor, but will still be edible; thus the "Never Refrigerate" stickers sometimes placed on tomatoes in supermarkets. Botanical description. Tomato plants are vines, initially, typically growing six feet or more above the ground if supported, although erect bush varieties have been bred, generally three feet tall or shorter. Indeterminate types are "tender" perennials, dying annually in temperate climates (they are originally native to tropical highlands), although they can live up to three years in a greenhouse in some cases. Determinate types are annual in all climates. Tomato plants are dicots, and grow as a series of branching stems, with a terminal bud at the tip that does the actual growing. When that tip eventually stops growing, whether because of pruning or flowering, lateral buds take over and grow into other, fully functional, vines. Tomato plant vines are typically pubescent, meaning covered with fine short hairs. These hairs facilitate the vining process, turning into roots wherever the plant is in contact with the ground and moisture, especially if there is some issue with the vine's contact to its original root. Most tomato plants have compound leaves, and are called regular leaf (RL) plants. But some cultivars have simple leaves known as potato leaf (PL) style because of their resemblance to that close cousin. Of regular leaves, there are variations, such as rugose leaves, which are deeply grooved, variegated, angora leaves, which have additional colors where a genetic mutation causes chlorophyll to be excluded from some portions of the leaves. Their flowers, appearing on the apical meristem, have the anthers fused along the edges, forming a column surrounded by the pistil's style. Flowers tend to be self-fertilizing. This is because they are native to the Americas, where there were no honeybees (which are native to the old world). Similarly, many plants of the Americas are self-fertilizing, while others are pollinated by flies, butterflies, moths, other insects, or other external forces that present in the Americas, that made it possible for some new world plants to originally require biotic pollination. Tomato fruit is classified as a berry. As a true fruit, it develops from the ovary of the plant after fertilization, its flesh comprising the pericarp walls. The fruit contains hollow spaces full of seeds and moisture, called locular cavities. These vary, among cultivated species, according to type. Some smaller varieties have two cavities, globe-shaped varieties typically have three to five, beefsteak tomatoes have a great number of smaller cavities, while paste tomatoes have very few, very small cavities. The seeds need to come from a mature fruit, and be dried fermented before germination. Botanical classification. In 1753 the tomato was placed in the genus "Solanum" by Linnaeus as "Solanum lycopersicum" L. (derivation, 'lyco', wolf, plus 'persicum', peach, "i.e.," "wolf-peach"). However, in 1768 Philip Miller placed it in its own genus, and he named it "Lycopersicon esculentum". This name came into wide use but was in breach of the plant naming rules. Technically, the combination "Lycopersicon lycopersicum" (L.) H.Karst. would be more correct, but this name (published in 1881) has hardly ever been used (except in seed catalogs, which frequently used it and still do). Therefore, it was decided to conserve the well-known "Lycopersicon esculentum", making this the correct name for the tomato when it is placed in the genus "Lycopersicon". However, genetic evidence ("e.g.," Peralta & Spooner 2001) has now shown that Linnaeus was correct in the placement of the tomato in the genus "Solanum", making the Linnaean name correct; if "Lycopersicon" is excluded from "Solanum", "Solanum" is left as a paraphyletic taxon. Despite this, it is likely that the exact taxonomic placement of the tomato will be controversial for some time to come, with both names found in the literature. Two of the major reasons that some still consider the genera separate are the leaf structure (tomato leaves are markedly different from any other "Solanum"), and the biochemistry (many of the alkaloids common to other "Solanum" species are conspicuously absent in the tomato). The tomato can with some difficulty be crossed with a few species of diploid Potato with viable offspring that are capable of reproducing. Such hybrids provide conclusive evidence of the close relationship between these genera. The Boyce Thompson Institute for Plant Research began sequencing the tomato genome in 2004 and is creating a database of genomic sequences and information on the tomato and related plants. A draft version of the full genome expected to be published by 2008. The genomes of its organelles (mitochondria and chloroplast) are also expected to be published as part of the project. Fruit or vegetable? Botanically, a tomato is the ovary, together with its seeds, of a flowering plant: therefore it is a fruit or, more precisely, a berry. However, the tomato is not as sweet as those foodstuffs usually called fruits and, from a culinary standpoint, it is typically served as part of a salad or main course of a meal, as are vegetables, rather than at dessert in the case of most fruits. As noted above, the term vegetable has no botanical meaning and is purely a culinary term. This argument has had legal implications in the United States. In 1887, U.S. tariff laws that imposed a duty on vegetables but not on fruits caused the tomato's status to become a matter of legal importance. The U.S. Supreme Court settled the controversy on May 10, 1893 by declaring that the tomato is a vegetable, based on the popular definition that classifies vegetables by use, that they are generally served with dinner and not dessert ("Nix v. Hedden" (149 U.S. 304)). The holding of the case applies only to the interpretation of the Tariff Act of March 3, 1883, and the court did not purport to reclassify the tomato for botanical or other purposes other than for paying a tax under a tariff act. Tomatoes have been designated the state vegetable of New Jersey. Arkansas took both sides by declaring the "South Arkansas Vine Ripe Pink Tomato" to be both the state fruit and the state vegetable in the same law, citing both its culinary and botanical classifications. In 2006, the Ohio House of Representatives passed a law that would have declared the tomato to be the official state fruit, but the bill died when the Ohio Senate failed to act on it. Tomato juice has been the official beverage of Ohio since 1965. A.W. Livingston, of Reynoldsburg, Ohio, played a large part in popularizing the tomato in the late 1800s. Due to the scientific definition of a fruit, the tomato remains a fruit when not dealing with US tariffs. Nor is it the only culinary vegetable that is a botanical fruit: eggplants, cucumbers, and squashes of all kinds (such as zucchini and pumpkins) share the same ambiguity. Pronunciation. The pronunciation of "tomato" differs in different English-speaking countries; the two most common variants are and. Speakers from the British Isles, most of the Commonwealth, and older generations among speakers of Southern American English typically say, while most American and Canadian speakers usually say. Many languages have a word that corresponds more to the former pronunciation, including the original Nahuatl word "tomato" from which they are all taken. The word's dual pronunciations were immortalized in Ira and George Gershwin's 1937 song "Let's Call the Whole Thing Off" ("You like and I like /You like and I like") and have become a symbol for nitpicking pronunciation disputes. In this capacity it has even become an American and British slang term: saying when presented with two choices can mean "What's the difference?" or "It's all the same to me." Safety. On October 30, 2006, the U.S. Centers for Disease Control and Prevention (CDC) announced that tomatoes might have been the source of a salmonella outbreak causing 172 illnesses in 18 states. The affected states included Arkansas, Connecticut, Georgia, Indiana, Kentucky, Maine, Massachusetts, Michigan, Minnesota, North Carolina, New Hampshire, Ohio, Pennsylvania, Rhode Island, Tennessee, Virginia, Vermont and Wisconsin. Tomatoes have been linked to seven salmonella outbreaks since 1990 (from the Food Safety Network). A 2008 salmonella outbreak caused the removal of tomatoes from stores and restaurants across the United States and parts of Canada. As of July 8, 2008, from April 10, 2008, the rare Saintpaul serotype of "Salmonella enterica" caused at least 1017 cases of salmonellosis food poisoning in 41 states throughout the United States, the District of Columbia, and Canada. As of July 2008, the U.S. Food and Drug Administration suspected that the contaminated food product was a common ingredient in fresh salsa, such as raw tomato, fresh jalapeño pepper, fresh serrano pepper, and fresh cilantro. It is the largest reported salmonellosis outbreak in the United States since 1985. New Mexico and Texas were proportionally the hardest hit by far, with 49.7 and 16.1 reported cases per million, respectively. The greatest number of reported cases occurred in Texas (384 reported cases), New Mexico (98), Illinois (100), and Arizona (49). There were at least 203 reported hospitalizations linked to the outbreak, it caused at least one death, and it may have been a contributing factor in at least one additional death. The CDC maintains that "it is likely many more illnesses have occurred than those reported." Applying a previous CDC estimated ratio of non-reported salmonellosis cases to reported cases (38.6:1), one would arrive at an estimated 40,273 illnesses from this outbreak. Tomato records. The heaviest tomato ever was one of 3.51 kg (7 lb 12 oz), of the cultivar 'Delicious', grown by Gordon Graham of Edmond, Oklahoma in 1986. The largest tomato plant grown was of the cultivar 'Sungold' and reached 19.8 m (65 ft) length, grown by Nutriculture Ltd (UK) of Mawdesley, Lancashire, UK, in 2000. The massive "tomato tree" growing inside the Walt Disney World Resort's experimental greenhouses in Lake Buena Vista, Florida may be the largest single tomato plant in the world. The plant has been recognized as a Guinness World Record Holder, with a harvest of more than 32,000 tomatoes and a total weight of 1,151.84 pounds (522 kg). It yields thousands of tomatoes at one time from a single vine. Yong Huang, Epcot's manager of agricultural science discovered the unique plant in Beijing, China. Huang brought its seeds to Epcot and created the specialized greenhouse for the fruit to grow. The vine grows golf ball-sized tomatoes which are served at Walt Disney World restaurants. The world record-setting tomato tree can be seen by guests along the Living With the Land boat ride at Epcot. On August 30, 2007, 40,000 Spaniards gathered in Buñol to throw of tomatoes at each other in the yearly Tomatina festival. Bare-chested tourists also included hundreds of British, French and Germans. Types. Varieties commonly grown by home gardeners include: Many varieties of processing tomatoes are grown commercially, but just five hybrid cultivars grown in California constitute over 60% of total production of processing tomatoes. Heritage and heirloom varieties with exceptional taste include: |